Encyclopaedia Britannica, 11th Edition, "Joints" to "Justinian I." / Volume 15, Slice 5
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JOINTS, in anatomy. The study of joints, or articulations, is known as Arthrology (Gr. ἄρθρον), and naturally begins with the definition of a joint. Anatomically the term is used for any connexion between two or more adjacent parts of the skeleton, whether they be bone or cartilage. Joints may be immovable, like those of the skull, or movable, like the knee.
Immovable joints, or synarthroses , are usually adaptations to growth rather than mobility, and are always between bones. When growth ceases the bones often unite, and the joint is then obliterated by a process known as synostosis , though whether the union of the bones is the cause or the effect of the stoppage of growth is obscure. Immovable joints never have a cavity between the two bones; there is simply a layer of the substance in which the bone has been laid down, and this remains unaltered. If the bone is being deposited in cartilage a layer of cartilage intervenes, and the joint is called synchondrosis (fig. 1), but if in membrane a thin layer of fibrous tissue persists, and the joint is then known as a suture (fig. 2). Good examples of synchondroses are the epiphysial lines which separate the epiphyses from the shafts of developing long bones, or the occipito-sphenoid synchondrosis in the base of the skull. Examples of sutures are plentiful in the vault of the skull, and are given special names, such as sutura dentata, s. serrata, s. squamosa, according to the plan of their outline. There are two kinds of fibrous synarthroses, which differ from sutures in that they do not synostose. One of these is a schindylesis , in which a thin plate of one bone is received into a slot in another, as in the joint between the sphenoid and vomer. The other is a peg and socket joint, or gomphosis , found where the fangs of the teeth fit into the alveoli or tooth sockets in the jaws.
Movable joints, or diarthroses , are divided into those in which there is much and little movement. When there is little movement the term half-joint or amphiarthrosis is used. The simplest kind of amphiarthrosis is that in which two bones are connected by bundles of fibrous tissue which pass at right angles from the one to the other; such a joint only differs from a suture in the fact that the intervening fibrous tissue is more plentiful and is organized into definite bundles, to which the name of interosseous ligaments is given, and also that it does not synostose when growth stops. A joint of this kind is called a syndesmosis , though probably the distinction is a very arbitrary one, and depends upon the amount of movement which is brought about by the muscles on the two bones. As an instance of this the inferior tibio-fibular joint of mammals may be cited. In man this is an excellent example of a syndesmosis, and there is only a slight play between the two bones. In the mouse there is no movement, and the two bones form a synchondrosis between them which speedily becomes a synostosis, while in many Marsupials there is free mobility between the tibia and fibula, and a definite synovial cavity is established. The other variety of amphiarthrosis or half-joint is the symphysis , which differs from the syndesmosis in having both bony surfaces lined with cartilage and between the two cartilages a layer of fibro-cartilage, the centre of which often softens and forms a small synovial cavity. Examples of this are the symphysis pubis, the mesosternal joint, and the joints between the bodies of the vertebrae (fig. 3).