PREFACE
I REGRET that pressure of other work has prevented the completion of this Volume within a reasonable time since the publication of Volume I. Had Volume II been written ten years ago, the discoveries made in the course of the last decade would have given an out-of-date character to much of the subject-matter. It is more especially in regard to the Ferns and the extinct members of the Gymnosperms that our outlook has been materially altered by recent contributions to Palaeobotany. It is, however, some satisfaction to be able to add that recent progress has been relatively slight in that part of the subject dealt with in the first volume.
The original intention was to complete the whole work in two volumes. Soon after the second volume was begun, it became evident that the remaining divisions of the plant-kingdom could not be included within the compass of a single volume. I decided, therefore, to take the consequences of having embarked on too ambitious a plan of treatment, and to preserve uniformity of proportion by reserving the seed-bearing plants for a third volume. The third volume will include the Pteridosperms, other than those briefly described in the final chapter of the present volume, and other classes of Gymnosperms. I propose also to devote such space as is available within the limits of a text-book to the neglected subject of the geographical distribution of plants at different stages in the history of the earth. It is my intention to complete Volume III with as little delay as possible. As I have written elsewhere, the past history of the Flowering plants needs special treatment, and anything more than a mere compilation can be adequately attempted only after considerable research and with the assistance of botanists possessing a special knowledge of different families of Angiosperms. The need of a critical examination of available data in regard to the geological history of this dominant group will not be lost sight of.
I am well aware that while certain genera have received an undue share of attention in the present volume, others have been ignored or treated with scant consideration. For this inconsistency I have no excuse to offer, beyond the statement that the subject is a large one, and selection is necessary even though the work consists of three volumes.
The publication in 1909 of a collection of excellent photographs of Palaeozoic Plants, with brief descriptive notes, by Mr Newell Arber, as one of a series of popular “Nature Books,” bears striking testimony to the remarkable spread of interest in the study of the vegetation of the past, which is one of the outstanding features in the recent history of botanical science.
In the list of illustrations I have mentioned the source of all figures which have been previously published. I would, however, supplement the statement of fact with an expression of thanks to corporate bodies and to individuals who have allowed me to make use of blocks, drawings, or photographs.
I wish to thank my colleague, Mr A. G. Tansley, for placing at my disposal several blocks originally published in the pages of the New Phytologist. To Professor Bertrand of Lille and to his son Dr Paul Bertrand I am indebted for several prints and descriptive notes of specimens in their possession. My friends Dr Nathorst of Stockholm and Dr Zeiller of Paris have generously responded to my requests for information on various points. I wish especially to thank Dr Kidston for several excellent prints of specimens in his collection and for the loan of sections. I have profited by more than one examination of his splendid collection at Stirling. Professor Weiss has generously allowed me to borrow sections from the Manchester University collections, more especially several which have been reproduced in the chapter devoted to the genus Lepidodendron. To Professor F. W. Oliver my thanks are due for the loan of sections from the collection under his charge at University College. I have pleasure also in thanking Dr Scott, not only for lending me sections of a Lepidodendron and for allowing me to use some drawings of Miadesmia originally made by Mrs Scott for reproduction in his invaluable book, Studies in Fossil Botany, but for kindly undertaking the laborious task of reading the proofs of this volume. It would be unfair to express my gratitude to Dr Scott for many helpful suggestions and criticisms, without explicitly stating that thanks to a friend for reading proofs must not be interpreted as an attempt to claim his support for all statements or views expressed. The General Editor of the Series, Mr A. E. Shipley, has also kindly read the proofs. I am under obligations also for assistance of various kinds to Prof. Thomas of Auckland, New Zealand, to Mr Boodle of Kew, to Mr D. M. S. Watson of Manchester, to Mr T. G. Hill of University College, and to Mr Gordon of Emmanuel College, Cambridge. I am indebted to the kind offices of Miss M. C. Knowles for the photograph of the specimen of Archaeopteris hibernica in the Irish National Museum, Dublin, reproduced on [page 561].
Many of the illustrations are reproduced from drawings by my wife: those made from the actual specimens are distinguished by the addition of the initials M. S. I am grateful to her also for some improvements in the letter-press. For the drawings made from sections and for some of the outline sketches I am responsible. I have availed myself freely of the facilities afforded by Professor McKenny Hughes in the Sedgwick Museum of Geology for the examination of specimens under the charge of Mr Newell Arber, the University Demonstrator in Palaeobotany. It is a pleasure to add that, as on former occasions, I am indebted to the vigilance of the Readers of the University Press for the detection of several errors which escaped my notice in the revision of the proofs.
A. C. SEWARD.
Botany School, Cambridge.
March 12, 1910.
| Fig. | Page | |
|---|---|---|
| [112.] | Sphenophyllostachys | 2 |
| [113.] | Sphenophyllostachys Römeri Sphenophyllum trichomatosum S. majus | 3 |
| [114, 115.] | Sphenophyllostachys fertilis [Council of the Royal Society of London.] | 4, 5 |
| [116.] | Sphenophyllostachys Dawsoni [Mr A. G. Tansley, Editor of the New Phytologist.] | 6 |
| [117.] | Cheirostrobus pettycurensis Pseudobornia ursina | 8 |
| [118.] | Psilotum triquetrum | 18 |
| [119.] | Psilotum triquetrum (anatomy) | 20 |
| [120.] | Tmesipteris tannensis | 22 |
| [121.] | Lycopodium (seven species) | 35 |
| [122.] | Lycopodium squarrosum | 36 |
| [123.] | Lycopodium cernuum | 37 |
| [124.] | Lycopodium obscurum | 38 |
| [125.] | Lycopodium (anatomy of stem) | 41 |
| [126.] | Lycopodium (anatomy of cones) | 45 |
| [127–129.] | Lycopodium cernuum (cone) [Council of the Royal Society of Edinburgh.] | 47–49 |
| [130.] | Selaginella grandis | 50 |
| [131.] | Selaginella (anatomy) | 52 |
| [132.] | Isoetes echinospora I. lacustris | 59 |
| [133.] | Isoetes lacustris (anatomy) | 62 |
| [134.] | Pleuromeia Sternbergi | 70 |
| [135.] | Selaginellites and Lycopodites | 80 |
| [136.] | Lycopodites lanceolatus [Council of the Geological Society of London.] | 81 |
| [137.] | Lycopodites falcatus | 83 |
| [138.] | Selaginellites primaevus | 86 |
| [139.] | Lycostrobus Scotti | 89 |
| [140.] | Picea excelsa | 94 |
| [141.] | Lepidodendron Sternbergii | 97 |
| [142.] | Sigillaria (leaves) | 98 |
| [143.] | Lepidodendron (leaves) | 99 |
| [144.] | Lepidodendron Veltheimianum | 101 |
| [145.] | Lepidodendron leaf-cushion | 102 |
| [146.] | Lepidodendron and Lepidophloios leaf-cushions | 104 |
| [147.] | Lepidophloios leaf-cushion | 108 |
| [148–155.] | Lepidodendron vasculare | 112–122 |
| [156.] | Knorria mirabilis | 125 |
| [157.] | Lepidodendron Veltheimianum (Ulodendron) | 129 |
| [158.] | Diagrammatic section illustrating the branch-theory of the Ulodendroid scar [Council of the Manchester Literary and Philosophical Society.] | 132 |
| [159.] | Pinus clausa | 134 |
| [160.] | Lepidophloios scoticus | 135 |
| [161.] | Halonia tortuosa | 136 |
| [162–167.] | Lepidodendron fuliginosum [Council of the Cambridge Philosophical Society.] | 143–147 |
| [168.] | Lepidodendron vasculare and L. fuliginosum | 148 |
| [169.] | Lepidodendron fuliginosum | 149 |
| [170–172.] | L. fuliginosum | 150–152 |
| [173.] | Lepidodendron obovatum | 154 |
| [174–176.] | Lepidodendron aculeatum [Oxford University Press: Annals of Botany.] | 155, 156 |
| [177.] | Stigmaria radiculosa | 157 |
| [178.] | Stigmarian rootlet | 158 |
| [179.] | Lepidodendron Harcourtii and L. fuliginosum | 162 |
| [180.] | Lepidodendron Wünschianum | 163 |
| [181, 182.] | L. Wünschianum | 165, 166 |
| [183, 184.] | L. Wünschianum [Editor of the New Phytologist.] | 168, 169 |
| [185.] | Lepidodendron Veltheimianum | 173 |
| [186.] | L. Veltheimianum and L. macrophyllum | 176 |
| [187.] | Lepidodendron australe [Dr H. Woodward, Editor of the Geological Magazine.] | 179 |
| [188, 189.] | Lepidostrobus | 183, 184 |
| [190.] | Lepidodendron and Lepidostrobi | 186 |
| [191.] | Lepidostrobus | 188 |
| [192.] | Spencerites insignis [Oxford University Press: Annals of Botany.] | 193 |
| [193.] | Sigillaria elegans, S. rugosa, S. tessellata, Omphalophloios anglicus | 197 |
| [194.] | Sigillaria McMurtriei | 199 |
| [195.] | Sigillaria mammillaris | 199 |
| [196.] | Sigillaria Brardi, S. laevigata, and Lepidodendron Wortheni | 200 |
| [197.] | Carica sp. | 202 |
| [198, 199.] | Sigillaria | 205, 206 |
| [200.] | Sigillaria Brardi | 212 |
| [201.] | Sigillariostrobus | 216 |
| [202.] | Sigillaria elegans and S. elongata | 220 |
| [203.] | Sigillaria Brardi | 225 |
| [204, 205.] | Stigmaria ficoides | 227, 228 |
| [206.] | Cyperus papyrus | 230 |
| [207, 208.] | Stages in the development of Sigillaria | 236 |
| [209.] | Stigmariopsis | 237 |
| [210.] | Stigmaria | 241 |
| [211.] | Bothrodendron punctatum | 250 |
| [212.] | Bothrodendron minutifolium, B. punctatum, B. kiltorkense and Lepidostrobus Olryi | 252 |
| [213.] | Bothrodendron minutifolium | 254 |
| [214.] | Bothrodendron Leslei [Trustees of the British Museum.] | 258 |
| [215.] | Bothrodendron mundum | 259 |
| [216.] | Bothrostrobus [Council of the Manchester Literary and Philosophical Society.] | 263 |
| [217.] | Omphalophloios | 265 |
| [218.] | Lepidocarpon Lomaxi | 273 |
| [219.] | Miadesmia and Bothrodendron | 276 |
| [220.] | Angiopteris evecta and Cycas revoluta | 283 |
| [221.] | Osmunda cinnamomea, O. regalis, and Todea barbara | 286 |
| [222.] | Schizaea elegans | 287 |
| [223.] | Aneimia rotundifolia | 288 |
| [224.] | Aneimia flexuosa, A. phyllitidis, Hymenophyllum, Matonia pectinata, Thyrsopteris elegans, Gleichenia | 289 |
| [225.] | Gleichenia dicarpa | 290 |
| [226.] | Gleichenites Rostafinskii, Gleichenia dicarpa, G. dichotoma | 290 |
| [227.] | Matonia pectinata [Council of the Royal Society.] | 292 |
| [228.] | Matonia pectinata | 293 |
| [229.] | Thyrsopteris elegans, Cyathea spinulosa, Dicksonia coniifolia, D. culcita, Davallia concinna, Alsophila excelsa | 294 |
| [230.] | Dicksonia Bertervana [Trustees of the British Museum.] | 295 |
| [231.] | Dipteris quinquefurcata, D. conjugata, D. Wallichii, and Polypodium quercifolium | 297 |
| [232.] | Davallia aculeata | 299 |
| [233.] | Polypodium Billardieri | 302 |
| [234.] | Polypodium quercifolium | 303 |
| [235.] | Hemitelia capensis | 304 |
| [236 a, 236 b.] | Pteris aquilina [Council of the Linnean Society of London.] | 305, 306 |
| [237.] | Matonia pectinata, Matonidium, Gleichenia dicarpa, and Trichomanes reniforme (anatomy) | 310 |
| [238.] | Trichomanes scandens [Editor of the New Phytologist.] | 311 |
| [239.] | Platyzoma microphylla [Editor of the New Phytologist.] | 312 |
| [240.] | Cyathea Imrayana [Editor of the New Phytologist.]. | 313 |
| [241.] | Angiopteris evecta and Marattia fraxinea | 317 |
| [242.] | Angiopteris evecta and Danaea | 318 |
| [243, 244.] | Angiopteris evecta | 319 |
| [245.] | Marattia fraxinea, M. Kaulfussii, Kaulfussia, and Marattiopsis Münsteri | 320 |
| [246.] | Ophioglossum vulgatum | 322 |
| [247.] | Botrychium virginianum | 322 |
| [248.] | Zalesskya gracilis | 327 |
| [249.] | Zalesskya diploxylon | 328 |
| [250.] | Thamnopteris Schlechtendalii | 330 |
| [251.] | Lonchopteris virginiensis | 331 |
| [252.] | Osmundites Dunlopi | 333 |
| [253, 254.] | Osmundites Kolbei [Editor of the Geological Magazine.] | 334, 335 |
| [255.] | O. Kolbei | 336 |
| [256.] | Cladophlebis denticulata, Todites Williamsoni, Discopteris Rallii, Kidstonia heracleensis, and Todeopsis primaeva | 340 |
| [257, 258.] | Cladophlebis denticulata | 342, 345 |
| [259.] | Klukia exilis [Council of the Cambridge Philosophical Society.] | 348 |
| [260.] | Ruffordia Goepperti | 349 |
| [261.] | Chrysodium lanzaeanum, Lygodium Kaulfussi, Marattia Hookeri | 350 |
| [262.] | Gleichenites longipennis, G. delicatula, G. Nordenskioldi and G. Zippei | 354 |
| [263.] | Gleichenites hantonensis [Council of the Palaeontographical Society.] | 356 |
| [264.] | Laccopteris elegans [Council of the Royal Society.] | 357 |
| [265.] | Matonidium Wiesneri, Marattiopsis marantacea, Gleichenites gracilis, Laccopteris Goepperti, and L. Muensteri | 358 |
| [266.] | Laccopteris polypodioides [Trustees of the British Museum.] | 359 |
| [267.] | Laccopteris [Trustees of the British Museum.] | 359 |
| [268.] | ? Laccopteris polypodioides [Trustees of the British Museum.] | 360 |
| [269.] | Matonidium Goepperti [Editor of the Encyclopaedia Britannica.] | 362 |
| [270.] | Senftenbergia elegans, Oligocarpia Brongniartii, Trichomanes sp., Hymenophyllum tunbridgense, Sphenopteris (Hymenophyllites) quadridactylites | 364 |
| [271.] | Coniopteris hymenophylloides [Council of the Manchester Literary and Philosophical Society.] | 368 |
| [272.] | C. hymenophylloides | 369 |
| [273.] | Coniopteris quinqueloba | 370 |
| [274.] | Coniopteris arguta | 371 |
| [275.] | Coniopteris arguta and C. hymenophylloides | 372 |
| [276.] | Oncopteris Nettvalli | 373 |
| [277.] | Protopteris punctata | 373 |
| [278.] | Laccopteris polypodioides, L. Muensteri, Dicksonia, Onychiopsis Mantelli, Hausmannia Sewardi, H. Kohlmanni, and Protopteris Witteana | 374 |
| [279.] | Adiantides antiquus and A. lindsayoides | 376 |
| [280.] | Onychiopsis Mantelli | 379 |
| [281.] | Dictyophyllum exile | 381 |
| [282.] | Dictyophyllum Nilssoni, Rhizomopteris Schenki, Camptopteris spiralis, and D. exile | 382 |
| [283.] | Dictyophyllum rugosum [Trustees of the British Museum.] | 384 |
| [284.] | Thaumatopteris Münsteri | 386 |
| [285.] | Clathropteris meniscoides | 387 |
| [286.] | Clathropteris egyptiaca [Editor of the Geological Magazine.] | 388 |
| [287.] | Camptopteris spiralis | 389 |
| [288.] | Hausmannia dichotoma | 391 |
| [289.] | Hausmannia sp. | 393 |
| [290.] | Alethopteris lonchitica, Lonchopteris rugosa, Sphenopteris Hoeninghausi, Parapecopteris neuropteroides,and Pecopteris (Dactylotheca) plumosa | 399 |
| [291.] | Ptychocarpus unita, Asterotheca Sternbergii, Danaeites sarepontanus, Hawlea Miltoni, H. pulcherrima, Scolecopteris elegans | 400 |
| [292.] | Dactylotheca plumosa | 405 |
| [293.] | D. plumosa | 406 |
| [294.] | Nathorstia angustifolia and N. latifolia | 410 |
| [295.] | Psaronius | 414 |
| [296.] | Psaronius infarctus, P. coalescens, P. musaeformis, and P. asterolithus | 416 |
| [297.] | Pecopteris Sterzeli | 419 |
| [298.] | Caulopteris peltigera and Megaphyton insigne | 421 |
| [299.] | Ptychopteris | 423 |
| [300.] | Dicksonia antarctica | 424 |
| [301.] | Rhacopteris sp. | 427 |
| [302.] | Noeggerathia foliosa | 429 |
| [303.] | Chiropteris Zeilleri [Annals of the South African Museum.] | 430 |
| [304.] | Tubicaulis solenites [Editor of the New Phytologist.] | 435 |
| [305.] | Botryopteris cylindrica | 439 |
| [306.] | Botryopteris ramosa | 441 |
| [307.] | Botryopteris antiqua | 442 |
| [308.] | Clepsydropsis antiqua, Etapteris Scotti, Diplolabis forensis, Zygopteris primaria, Stauropteris oldhamia | 444 |
| [309.] | Diplolabis forensis, Botryopteris forensis, Corynepteris coralloides, Schizopteris pinnata | 445 |
| [310.] | Metaclepsydropsis duplex, Stauropteris oldhamia, Ankyropteris scandens | 450 |
| [311.] | Ankyropteris Grayi | 451 |
| [312.] | Thamnopteris Schlechtendalii, Ankyropteris corrugata, A. bibractensis | 453 |
| [313.] | Ankyropteris bibractensis | 454 |
| [314.] | Ankyropteris corrugata | 457 |
| [315.] | Ankyropteris corrugata [Editor of the New Phytologist.] | 458 |
| [316, 317.] | Ankyropteris corrugata | 459, 460 |
| [318.] | Etapteris Scotti [Editor of the New Phytologist.] | 462 |
| [319.] | Etapteris, Botryopteris forensis | 463 |
| [320.] | Stauropteris oldhamia [Editor of the New Phytologist.] | 464 |
| [321.] | Stauropteris oldhamia | 467 |
| [322.] | Stauropteris oldhamia [Editor of the New Phytologist.] | 468 |
| [323.] | Stauropteris [Editor of the New Phytologist.] | 469 |
| [324.] | Asterochlaena laxa [Editor of the New Phytologist.] | 472 |
| [325.] | Sporocarp-like bodies (? Sagenopteris) | 478 |
| [326.] | Regnellidium diphyllum, Sagenopteris rhoifolia | 479 |
| [327.] | Sagenopteris Phillipsi [Trustees of the British Museum.] | 480 |
| [328.] | Sagenopteris Phillipsi [Council of the Manchester Literary and Philosophical Society.] | 481 |
| [329.] | Taeniopteris multinervis, Lesleya Delafondi | 487 |
| [330.] | Taeniopteris Carnoti, T. spatulata, T. coriacea | 490 |
| [331.] | Taeniopteris Carruthersi [Annals of the South African Museum.] | 491 |
| [332.] | Taeniopteris vittata | 493 |
| [333.] | Weichselia Mantelli, W. erratica | 495 |
| [334.] | Glossopteris Browniana [Council of the Geological Society of London.] | 499 |
| [335, 336.] | Glossopteris Browniana [Trustees of the British Museum.] | 500, 501 |
| [337.] | Vertebraria indica | 502 |
| [338.] | Vertebraria indica, Onoclea struthiopteris | 503 |
| [339.] | Glossopteris fronds attached to rhizome | 504 |
| [340, 341.] | Glossopteris indica, G. angustifolia [Trustees of the British Museum.] | 506, 507 |
| [342.] | Glossopteris angustifolia var. taeniopteroides [Council of the Geological Society.] | 508 |
| [343.] | Blechnoxylon talbragarense | 509 |
| [344.] | Glossopteris retifera [Trustees of the British Museum.] | 511 |
| [345.] | Gangamopteris cyclopteroides [Trustees of the British Museum.] | 515 |
| [346.] | Arberia sp. | 517 |
| [347.] | Lesleya simplicinervis | 518 |
| [348.] | Neuropteridium validum [Trustees of the British Museum.] | 520 |
| [349.] | Neuropteridium intermedium | 522 |
| [350.] | Cardiopteris frondosa | 524 |
| [351.] | Gunnera manicata | 527 |
| [352.] | Sphenopteris obtusiloba, Pecopteris arborescens, Sphenopteris furcata | 529 |
| [353.] | Sphenopteris affinis | 531 |
| [354.] | Palmatopteris, Mariopteris, Diplotmema Zeilleri, Neuropteris macrophylla, N. heterophylla, N. Scheuchzeri, Alloiopteris Essinghii | 535 |
| [355.] | Cephalotheca mirabilis | 536 |
| [356.] | Thinnfeldia odontopteroides, Ptilozamites [Council of the Geological Society.] | 539 |
| [357.] | Thinnfeldia odontopteroides [Council of the Geological Society.] | 540 |
| [358.] | Thinnfeldia odontopteroides [Annals of the South African Museum.] | 541 |
| [359.] | Thinnfeldia rhomboidalis | 542 |
| [360.] | Lomatopteris jurensis, L. Schimperi, Thinnfeldia rhomboidalis | 544 |
| [361.] | Ptilozamites Heeri | 547 |
| [362.] | Ctenopteris cycadea | 549 |
| [363.] | Dichopteris visianica | 551 |
| [364.] | Alethopteris lonchitica, Mariopteris muricata, Odontopteris cf. alpina | 553 |
| [365.] | Odontopteris minor | 554 |
| [366.] | Odontopteris genuina, Callipteridium gigas, Callipteris Pellati, C. lyratifolia | 557 |
| [367.] | Callipteris conferta | 559 |
| [368.] | Archaeopteris hibernica | 561 |
| [369.] | Archaeopteris hibernica, A. archetypus, A.fissilis, A. fimbriata | 564 |
| [370.] | Neuropteris with Cyclopteris leaflets [From a block received from Mr Carruthers.] | 566 |
| [371.] | Neuropteris heterophylla | 568 |
| [372.] | Neuropteris macrophylla | 569 |
| [373.] | Neuropteris Scheuchzeri | 570 |
| [374.] | Linopteris neuropteroides | 573 |
| [375.] | Alethopteris Serlii | 575 |
| [376.] | Pecopteris arborescens | 578 |