The Oral Cavity

The mouth in the Crocodilia is large, as is well known, and may be opened very wide. It is bounded anteriorly and laterally by the teeth of the two jaws; these teeth were described in connection with the skull. The mucous membrane of the roof of the mouth and of the dorsum of the tongue, especially the former, exhibits numerous small papillæ (see [page 160]), and among these, in the posterior region of the mouth, are the ducts of mucous glands.

Fig. 33. Interior of the Mouth of A. Mississippiensis

f, transverse fold at the base of the tongue; v, v, velum palatinum.

The tongue extends from just back of the mandibular symphysis to the glottis. It is attached throughout its entire ventral side except for a short distance at its tip, so that it may be elevated and depressed but not protruded. Among the papillæ on its dorsal surface are sense organs, said to be tactile and gustatory corpuscles (see page 165). The posterior margin of the tongue is elevated as a transverse fold that meets a corresponding fold, the velum palatinum, from the lower side of the palate and completely shuts off the mouth from the openings of the trachea and gullet ([Fig. 33]). Into this hinder chamber open the posterior nares, so that the animal can open its mouth under water without getting water into its trachea; or it may, while holding its prey in its mouth, come to the surface to breathe, without danger of letting water into its trachea. The nasal passages, leading from the nostrils to the posterior nares, are, of course, completely inclosed by bone, as described in connection with the skull. Ventral to the larynx and posterior part of the mouth is the large, shield-shaped hyoid apparatus, [Fig. 25], h, also described in connection with the skull.