CHAPTER XXXVIII.
PHYSIOLOGY OF THE NERVOUS SYSTEM.
768. The brain is regarded by physiologists and philosophers as the organ of the mind. Most writers consider it as an aggregate of parts, each charged with specific functions, and that these functions are the highest and most important in the animal economy. To the large brain, or cerebral lobes, they ascribe the seat of the faculties of thinking, memory, and the will. In man, this lobe extends so far backward as to cover the whole of the cerebellum. To the cerebellum, or little brain, is ascribed the seat of the animal, or lower propensities.
769. “The constant relation between mental power and development of brain, explains why capacities and dispositions are so different. In infancy, for example, the intellectual powers are feeble and inactive. This arises partly from the inaptitude of a still imperfect brain; but in proportion as the latter advances toward its mature state, the mental faculties also become vigorous and active.”
770. We are able, in most instances, at least, to trace a correspondence between the development of the cerebral lobes and the amount of intelligence possessed by the person. The weight of the brain in man to that of the whole body varies 347 in different individuals. The heaviest brain on record was that of Cuvier, which weighed 4 pounds and 13½ ounces.
768–772. Give the physiology of the nervous system. 768. How is the brain regarded by physiologists and philosophers? What do they ascribe to the cerebrum? To the cerebellum? 769. What does the relation between mental powers and development of brain explain? 770. What is said respecting the correspondence between the development of the brain and the amount of intelligence possessed by the person? What is said of the weight of the brain?
771. The brain likewise holds an important relation to all the other organs of the system. To the muscular system it imparts an influence which induces contraction of the fibres. By this relation they are brought under the control of the will. To the skin, eye, and ear, it imparts an influence that gives sensibility, or the power of feeling, seeing, hearing, &c.
772. Again, the involuntary functions of the different portions of the system are more or less influenced by the brain. If the action of this central organ of the nervous system is destroyed, the functions of the digestive, respiratory, and circulatory apparatuses will be much disturbed or entirely suppressed.
773. The brain is the seat of sensation. It receives the impressions made on all parts of the body, through the medium of the sensitive nerves. That the impressions of external objects, made on these nerves, be communicated to the brain, where sensation is perceived, it is necessary that they be not diseased or injured.
Observation. There is a plain distinction between sensations and impressions; the latter are the changes produced in the extremities of the nerve; the former, the changes produced in the brain and communicated to the mind.
774. What part of the brain receives the impressions or has the most intimate relation with the intellectual faculties is unknown. Some portions, however, are of less importance than others. Large portions of the cortical, or outer part, are frequently removed without affecting the functions of this 348 organ. Pieces of the medullary, or central parts, have been removed by injuries without impairing the intellect or destroying life. This organ, although it takes cognizance of every sensation, is, of itself, but slightly sensible. It may be cut, or parts may be removed without pain, and the individual, at the same times retain his consciousness.
771. What is said of the relation of the brain to all of the organs of the body? 772. Are the involuntary functions of different parts of the system influenced by the brain? 773. Where is sensation perceived? By what agency are the impressions of external objects conveyed to the brain? What is the difference between sensations and impressions? 774. Is it known what part of the brain has the most intimate relation with the intellectual faculties?
775. The brain is the seat of the will. It superintends the physical as well as the mental movements, and the medium of communication from this organ to the muscles, or the parts to be moved, is the motor nerves. If the brain is in a quiescent state, the muscles are at rest; if, by an act of the will, the brain sends a portion of nervous influence to a voluntary muscle, it immediately contracts, and those parts to which the muscle is attached move. There is no perceptible interval between the act of the will and the motion of the part.
776. Some physiologists assert, that the medulla oblongata is the point at which excitement to motion commences, and sensation terminates; and also, that it possesses the power of originating motion in itself.
Observation. The medulla oblongata, unlike the brain, is highly sensitive; if slightly punctured, convulsions follow; if much injured, respiration, or breathing, immediately ceases.
777. It is remarkable that the nerves which arise from the right side of the spinal cord communicate with the left hemisphere of the cerebrum, and vice versa; this results from the crossing of the fibres in the medulla oblongata. It follows from this, that if the right side of the brain receives an injury, the parts of the opposite side of the body lose their sensibility and motion.
Observations. 1st. If the cranial nerves which are connected 349 by a single root are divided, only the sensation of the part to which they are distributed is lost. Thus, if the optic nerve is divided, the sense of vision disappears, but the motions of the eye are performed as readily as before. But, if the spinal nerves are divided, both sensation and motion of the part to which they lead are destroyed.
What portions have been removed without impairing the intellect? What is remarkable of the brain? 775. What is the influence of the brain upon the muscles? 776. What do some physiologists assert of the medulla oblongata? 777. What is remarkable of the nerves? Give the 1st observation relative to the cranial nerves.
2d. When the spinal cord is divided or compressed, as in fractures of the spinal column, all parts below the fracture are paralyzed, though the nerves leading to these parts may be uninjured.
3d. Again, one side of the body or one limb may become insensible, and the power to move it, be perfectly retained; or the reverse of this may happen—the power of motion will be lost while sensation remains. In the former instance, the function of the posterior, or sensitive column of the spinal cord on one side is destroyed; in the latter, the anterior, or motor column is affected.
4th. In some cases, both sensation and motion of one side of the body or one limb are destroyed. In such instances, both the anterior and the posterior columns of one side of the spinal cord are diseased.
778. Vigorous and controllable muscular contraction requires a sound and well-developed brain. If this organ is defective in these particulars, the movements will be inefficient, and may be irregular. The central organ of the nervous system must, likewise, be in an active condition, to induce regular, steady, and controllable muscular movements.
Observations. 1st. Persons who have suffered from apoplexy and other severe diseases of the brain, have an involuntary trembling of the limbs, which results from a weakened state of the nervous system.
To the spinal nerves. What is said of the compression of the spinal cord? Give the 3d observation relative to the spinal nerves. The 4th observation. 778. Upon what does vigorous controllable muscular contraction depend? What causes the involuntary trembling of the limbs in persons who have suffered from apoplexy?
2d. The tremor of the hand, that lessens the usefulness or incapacitates the fine artist or skilful mechanic, in the prime of life, from pursuing their vocations, may be, and is often, induced by the influence of intoxicating drink, which debilitates and disorganizes the brain.
3d. The tottering step, trembling hand, and shaking head of the aged invalid, are the results of diminished nervous energy, so that steady muscular contraction, so essential to regular movements, cannot be maintained.
779. No difference can be discovered in the structure of the several kinds of nerves in any part of their course, and the functions they are designed to perform can only be known by ascertaining the place of their origin. The nerves may be functionally divided into five groups.
780. 1st. Nerves of special sensation. These are the first, second, eighth, and it may be one of the branches of the fifth pair of cranial nerves. The function of these nerves is particularly described in the chapters upon the senses of smell, vision, hearing, and taste.
781. 2d. Nerves of general sensation. These embrace the fifth pair of cranial nerves, and the thirty-one pairs of spinal nerves. In those parts that require sensation for their safety and the performance of their functions, there is an abundant supply of sensitive nervous filaments. The nerves of sensation are mostly distributed upon the skin. Few filaments ramify upon the mucous membranes and muscles.
Observations. 1st. The painful sensations experienced in the face, and in the teeth or jaws, (tic douloureux and toothache,) are induced by irritation and disease of a portion of the filaments of the fifth pair of cranial nerves.
The tremor of the hand among some mechanics in the prime of life? The tottering step of the aged invalid? 779. What is said relative to the structure of the nerves? How may they be divided? 780. Give the nerves of special sense. 781. Those of general sensation. Where are the nerves of sensation distributed? What causes tic douloureux?
2d. The unpleasant sensation sometimes experienced when we hear the grating of a file or saw, is produced by the connection of the nerve that passes across the drum of the ear with the fifth cranial nerve.
3d. When pressure is made on the trunk of a nerve, the sensibility of the part where the nerve ramifies is modified. This is illustrated, when pressure is made upon the large nerve of the lower extremity (sciatic) in sitting upon a hard bench. The foot is then said to be “asleep.”
4th. When the trunk of a nerve is diseased or injured, the pain is experienced in the outer extremity of the nerve. A blow upon the elbow, which causes a peculiar sensation in the little finger and one side of the ring finger, affords a familiar illustration. This sensation is produced by injuring the ulnar nerve, which is distributed to the little finger.
782. 3d. Nerves of motion. These are the third, sixth, and twelfth pairs of cranial nerves, and the thirty-one pairs of spinal nerves. These nerves are distributed to the fibres of the five hundred muscles of the body. The functions of the muscular are different from those of the sensitive nerves. The former are provided for the purpose of motion, and not of feeling. Hence, muscles may be cut, and the pain will be slight, compared with the cutting of the skin. This may be called muscular pain. Weariness is a sensation recognized by one set of muscular nerves.
783. So uniformly is a separate instrument provided for every additional function, that there is strong reason to regard the muscular nerves, although running in one sheath, as in reality double, and performing distinct functions. Sir Charles Bell, in his work on the Nervous System, endeavors to show, 352 that one set of nervous fibres conveys the mandate from the brain to the muscle, and excites the contraction; and that another set conveys, from the muscle to the brain, a peculiar sense of the state or degree of contraction of the muscle, by which we are enabled to judge of the amount of stimulus necessary to accomplish the end desired. This is obviously an indispensable piece of information to the mind in regulating the movements of the body.
How is the peculiar sensation accounted for when we hear the grating of a file or saw? What produces the sensation when the foot is said to be “asleep?” What is the effect when the ulnar nerve is injured by a blow? 782. Give the nerves of motion. What is said of the functions of the muscular nerves? 783. What does Sir Charles Bell endeavor to show?
784. 4th. Nerves of respiration. These are the fourth, seventh, ninth, tenth, and eleventh pair of cranial nerves, also the phrenic and the external respiratory nerve. All of these nerves have their origin in a distinct tract or column, called the lateral, in the upper part of the spinal cord. Hence it is sometimes named the respiratory column. These nerves are distributed to one of the muscles of the eye; to the muscles of the face; to the tongue, pharynx, œsophagus, stomach, heart, lungs, diaphragm, and some of the muscles of the neck and chest.
785. It is through the instrumentality of the accessory, phrenic, and external respiratory nerves, (10, 11, 12, 13, fig. 132,) that the muscles employed in respiration are brought into action without the necessity of the interference of the mind. Though to a certain extent they may be under the influence of the will, yet it is only in a secondary degree. No one can long suspend the movements of respiration;[20] for in a short time, instinctive feeling issues its irresistible mandate, which neither requires the aid of erring wisdom, nor brooks the capricious interference of the will.
Fig. 132.
Fig. 132. The distribution of the respiratory nerves. a, Section of the brain and medulla oblongata. b, The lateral columns of the spinal cord. c, c, The respiratory tract of the spinal cord. d, The tongue. e, The larynx. f, The bronchia. g, The œsophagus. h, The stomach. i, The diaphragm. 1, The pneumogastric nerve. 2, The superior laryngeal nerve. 3, The recurrent laryngeal nerve. (These two ramify on the larynx.) 4, The pulmonary plexus of the tenth nerve. 5, The cardiac plexus of the tenth nerve. These two plexuses supply the heart and lungs with nervous filaments. 7, The origin of the fourth pair of nerves, that passes to the superior oblique muscle of the eye. 8, The origin of the facial nerve, that is spread out on the side of the face and nose. 9, The origin of the glosso-pharyngeal nerve, that passes to the tongue and pharynx. 10, The origin of the spinal accessory nerve. 11, This nerve penetrating the sterno-mastoideus muscle. 12, The origin of the internal respiratory or phrenic nerve, that is seen to ramify on the diaphragm. 13, The origin of the external respiratory nerve, that ramifies on the pectoral and scaleni muscles.
784. Give the respiratory nerves. What is said in reference to the respiratory nerves? 785. Through the agency of what nerves are the respiratory muscles brought into action? Explain fig. 132. 354
786. The fourth, seventh, and tenth pairs of nerves, (7, 8, 9, fig. 132,) with the spinal accessory, phrenic, and external respiratory, are not only connected with the function of respiration, but contribute to the expression of the passions and emotions of the mind.
787. The influence of this order of nerves in the expression of the passions, is strikingly depicted in Sir Charles Bell’s Treatise on the Nervous System. “In terror,” he remarks, “we can readily conceive why a man stands with his eyes intently fixed on the object of his fears—the eyebrows elevated, and the eyeballs largely uncovered; or why, with hesitating and bewildered steps, his eyes are rapidly and wildly in search of something. In this way, we only perceive the intense application of his mind to the objects of his apprehension, and its direct influence on the outward organs.”
Can respiration be suspended for any considerable length of time? 786. What nerves contribute to the expression of the passions and emotions of the mind? 787, 788. What does Sir Charles Bell say of the influence of this order of nerves in the expression of the passions?
788. “But when we observe him further, there is a spasm in his breast; he cannot breathe freely; the chest remains elevated, and his respiration is short and rapid. There is a 355 gasping and convulsive motion of his lips, a tremor on his hollow cheeks, a gasping and catching of his throat; his heart knocks at his ribs, while yet there is no force in the circulation—the lips and cheeks being ashy pale.”
789. “These nerves are the instruments of expression, from the smile upon the infant’s cheek, to the last agony of life. It is when the strong man is subdued by this mysterious influence of soul on body, and when the passions may be truly said to tear the heart, that we have the most afflicting picture of human frailty, and the most unequivocal proof that it is the order of functions we have been considering, that is thus affected. In the first struggle of the infant to draw breath, in the man recovering from a state of suffocation, and in the agony of passion, when the breast labors from the influence at the heart, the same system of parts is affected, the same nerves, the same muscles, and the symptoms or character have a strict resemblance.”
790. The seventh pair of nerves not only communicates the purposes of the will to the muscles of the face, but at the same time it calls them into action, under the influence of instinct and sympathy. On this subject a late writer remarks, “How expressive is the face of man! How clearly it announces the thoughts and sentiments of the mind! How well depicted are the passions on his countenance! tumultuous rage, abject fear, devoted love, envy, hatred, grief, and every other emotion, in all their shades and diversities, are imprinted there, in characters so clear that he that runs may read! How difficult, nay, how impossible, is it to hide or falsify the expressions which indicate the internal feelings! Thus conscious guilt shrinks from detection, innocence declares its confidence, and hope anticipates with bright expectation.”
Observation. The fifth pair of nerves (fig. 126) is distributed 356 to the parts of the face on which the seventh pair ramifies. The former serves for sensation, the latter for motion. Thus, when the seventh pair of nerves is divided, or its functions destroyed by disease, the side affected loses all power of expression, though sensation remains unaffected. On the contrary, if we divide the fifth pair, sensation is entirely destroyed, while expression remains.
789. Are they also the instruments of expression, either of joy or grief? 790. What is said in reference to the seventh pair of nerves? Where is the fifth pair of nerves distributed?
791. 5th. The sympathetic nerve. This nerve confers vitality on all the important portions of the system. It exerts a controlling influence over the involuntary functions of digestion, absorption, secretion, circulation, and nutrition. Every portion of the body is, to a certain extent, under its influence, as filaments from this system of nerves accompany the blood-vessels throughout their course.
792. An important use of the sympathetic nerve is to form a communication of one part of the system with another, so that one organ can take cognizance of the condition of every other, and act accordingly. If, for example, disease seizes the brain, the stomach, by its sympathetic connection, knows it; and as nourishment would add to the disease, it refuses to receive food, and perhaps throws off what has already been taken. Loss of appetite in sickness is thus a kind provision of nature, to prevent our taking food when it would be injurious; and following this intimation, we, as a general rule, should abstain from food until the appetite returns.
Fig. 133.
Fig. 133. A back view of the brain and spinal cord. 1, The cerebrum. 2, The cerebellum. 3, The spinal cord. 4, Nerves of the face. 5, The brachial plexus of nerves. 6, 7, 8, 9, Nerves of the arm. 10, Nerves that pass under the ribs, 11, The lumbar plexus of nerves. 12, The sacral plexus of nerves 13, 14, 15, 16, Nerves of the lower limbs.
What is the function of this nerve? What is the effect if the seventh pair is divided, or its function destroyed by disease? 791. What is said of the sympathetic nerve? 792. What is the use of the sympathetic system? Explain fig. 133.
Note. Let the anatomy and physiology of the nervous system be reviewed from figs. 131, 132, 133, or from anatomical outline plate. No. 8.