STARFISHES, SEA URCHINS, AND TREPANGS
We have now arrived at the point (phylum Echinodermata, "spiny-skinned") where a distinctly new type of interior structure appears in the possession by animals of a hollow space (cœlom) between the outer skin and the wall of the digestive tube which now becomes occupied by definite organs instead of by an almost uniform mesenchyme, as in the sponges and cœlenterates. These organs arise from an interior lining membrane called "mesoderm."
| Photo, American Museum of Natural History |
| STARFISH AND OTHER TYPICAL LIFE IN A TIDE POOL |
Henceforth, therefore, we shall deal with cœlomate animals, among which the echinoderms are lowest in rank. The simplest of them is the "sea lily" which lives rooted on the bottom in deep water, and sways about on a slender, jointed stalk, looking much like the flower after which it is named. It is of interest chiefly as a survivor of the tribe of crinoids that were so varied and numerous in early Paleozic times that massive Devonian limestones are composed largely of their remains; and the type has changed little through the ages. It consists typically of a cup, mounted on its stem like the calyx of a flower, and composed of circles of calcareous plates, definite in form and in relative position, that contain and protect a well-organized body. Surrounding the open mouth of the cup is a circle of long, jointed, much-branched tentacles that sweep the water, capture passing prey, and bring it into the mouth of the crinoid within the circling base of the arms.
| Photo, American Museum of Natural History |
| CORAL FORMATIONS OF THE GREAT BARRIER REEF, AUSTRALIA |
If now you were to cut off its stalk, lay the crinoid on the sand, mouth down and arms outspread, beside a brittle star or a basket fish, which also have many-branched arms, it might be difficult to tell them apart, yet they represent different orders; and from this, by way of the naked serpent star, it is but a short transition to the starfish, where the arms are no longer tentaclelike, but are simply pointed extensions of a central body; this, in fact, is the case, for now they are no longer prehensile organs, but are supports, mainly serviceable in locomotion, and the stomach and ovaries are partly lodged in them. The main point just now, however, is the fact that here, and in the successive changes of form to be shown, the pattern of plates that form a strengthening mosaic in the skin of the central part of the body remains identical.
All starfishes are not as prettily symmetric as our familiar five-finger. Some are shorter in the arms, and much broader and thicker in the body; and if you will examine a collection of preserved specimens of the echinoderms you will see that you can trace gradation of form right around to the bun-shaped cake urchin, on whose top the five-pointed star is printed, and thence to the globular sea egg, which the French called "sea urchin," using one of their names for the hedgehog. Furthermore, the five sections of the shell of the urchin, which represent the five arms of the starfish folded forward and grown together into a spherical case, are to be traced again, outlined by appendages, in the elongated and leathery hide of the trepangs and sea cucumbers of the order Holothuria.
It is as an illustration of homology, that is, the resemblance between parts that have the same relation to the typical plan of structure, and as an example of how almost endless variations of form may arise within a single type, that the echinoderms are of most interest. Otherwise it may be said that they serve as food for fishes and some other creatures, including coastwise savages, and as curiosities in geological museums and in aquaria; and that starfishes are sadly destructive of cultivated oyster beds. We may therefore dismiss them, and devote a page or two to the worms.