Mackenzie Valley—Great Lakes—Mississippi Valley route and tributaries

Easily the longest route of any in the Western Hemisphere is that extending from the Mackenzie Valley past the Great Lakes and down the Mississippi River, including its tributaries. Its northern terminus is on the Arctic coast in the regions of Kotzebue Sound, Alaska, and the mouth of the Mackenzie River, while its southern end lies in Patagonia. During the spring migration some of the shore birds traverse the full extent of this great path, and it seems likely that the nighthawk, the barn swallow, the blackpolled warbler, and individuals of several other species that breed north to Yukon and Alaska must twice each year cover the larger part of it.

For more than 3,000 miles—from the mouth of the Mackenzie to the delta of the Mississippi—this route is uninterrupted by mountains. In fact, there is not even a ridge of hills high enough to interfere with the movements of the feathered travelers, and the greatest elevation above sea level is less than 2,000 feet. Well timbered and watered, the entire region affords ideal conditions for the support of its great hosts of migrating birds. It is followed by such vast numbers of ducks, geese, shore birds, blackbirds, sparrows, warblers, and thrushes, that observers stationed at favorable points in the Mississippi Valley daring the height of migration can see a greater number of species and individuals than can be noted anywhere else in the world.

Starting in the region of Kotzebue Sound, Alaska, the route extends eastward across northern Alaska and joins another that has its origin at the mouth of the Mackenzie River. The line of flight then trends a little east of south through the great lake system of central Canada, where it is joined by two or three other routes from the northeast that have their origin on the central Arctic coast. Continuing southward the migrating flocks are constantly augmented by additions to their numbers as they pass over the great breeding grounds of central and southern Canada. Upon reaching the headwaters of the Missouri and Mississippi Rivers the route follows these streams to the Gulf coast, Arriving in this latitude many species, including the ducks and geese, the robin, the myrtle warbler, and some others spread out east and west for their winter sojourn. Others, despite the perils of a trip involving a flight of several hundred miles across the Gulf of Mexico, strike out boldly for Central America and South America. This part of the route is a broad "boulevard" extending from northwestern Florida to eastern Texas and reaching southward across the Gulf of Mexico to Yucatan and the Isthmus of Tehuantepec ([fig. 17], route 4). In other words, when most of the birds (chiefly shore birds and land birds) that have come south directly through the Mississippi Valley have reached the coastal plains of Mississippi and Louisiana, they continue directly across the Gulf of Mexico. This route appears to have preference over the safer but more circuitous land or island routes by way of Texas and Florida. During the height of migration some of the islands off the coast of Louisiana are wonderful observation points for the student of birds, as the feathered travelers literally swarm over them.

Present detailed knowledge of the chief tributaries to the Mackenzie—Great Lakes—Mississippi Valley highway relates chiefly to waterfowl. Reference already has been made to the flight of the black ducks (p. 58) that reach the Mississippi Valley from southern Ontario. Some individuals of this species banded at Lake Scugog, Ontario, have been recaptured in succeeding seasons in Wisconsin and Manitoba, but the majority have been retaken at points south of the junction of the Ohio River with the Mississippi, definitely indicating their route of travel from southern Ontario.

A second route that joins the main artery on its eastern side is the one used by the blue goose, the migration route of which is probably more nearly due north and south than that of any other North American bird. The breeding grounds, which only in recent years have been discovered, are mainly in the Fox Basin region of Baffin Island and on Southampton Island. In fall these geese work southward, chiefly along the eastern shore of Hudson Bay, and upon reaching the southern extremity of James Bay they take off for what is practically a nonstop flight to the great coastal marshes of Louisiana west of the delta of the Mississippi River. In some seasons the flocks make intermediate stops among the islands and sand bars of the Mississippi, as they are occasionally common in the general vicinity of Memphis, Tenn. Most of the birds push on, however, and during the period from the first of November to the last of March fully 90 percent of the species are concentrated in the area between the Sabine and the Mississippi Rivers. On the return trip northward there is sometimes a tendency for some of the blue geese to veer off toward the northwest, as they are occasionally abundant in eastern South Dakota and southeastern Manitoba. It is of particular interest to note that while some other geese and many ducks start their northward journey at the first sign of awakening spring, the blue goose remains in its winter quarters until the season there is far advanced, seemingly aware that its own breeding grounds in the Arctic are still in the grip of winter.