Movements of species and groups
Some species begin their fall migrations early in July and in some parts of the country distinct southward movements can be detected from then until the beginning or middle of winter. For example, many shore birds start south in the early part of July, while the goshawks, snowy owls, redpolls, Bohemian waxwings, and many others do not leave the North until forced to do so by the advent of severe winter weather, or by lack of the customary food. Thus an observer in the northern part of the United States may record an almost unbroken southward procession of birds from midsummer to winter, and note some of the returning migrants as early as the middle of February. While on their way north, purple martins have been known to arrive in Florida late in January and, among late arrivals, the northern movement may continue into the first week of June. In some species the migration is so prolonged that the first arrivals in the southern part of the breeding range will have performed their parental duties while others of that species are still on their way north.
A study of these facts indicates that sometimes there exists a very definite relationship between what we may term northern and southern groups of individuals of the same species. A supposition is that for a species with an extensive latitudinal breeding range, and which has a normal migration, those individuals that nest farthest south migrate first and proceed to the southern part of the winter range; those that occupy the central parts of the breeding range migrate next and travel to regions in the winter range north of those occupied by the first group; and finally the individuals breeding farthest north are the last to start their autumn migration and they remain farthest north during the winter. In other words, this theory supposes that the southward movement of the species is such that the different groups maintain their relative latitudinal position with each other. The black and white warbler furnishes an example. The breeding range of this bird extends west and northwest from northern Georgia and South Carolina to New Brunswick, extending also in a western and northwestern direction as far as Great Bear Lake in northwestern Canada ([fig. 1]). It spends the winter in southern Florida, the West Indies, central Mexico, Central America, and northwestern South America. In the southern part of its breeding range it is nesting in April, but those that summer in New Brunswick do not reach their nesting grounds before the middle of May. Therefore, about 50 days are required for these northbound birds to cross the breeding range, and if 60 days be allowed for nest building, egg laying, incubation, care of young, and molt, they would not be ready to start southward before the middle of July ([fig. 2.]). Then another 50-day trip south, and the earliest migrants from the northern areas would reach the Gulf Coast in September. But both adults and young have been observed at Key West, Fla., by the middle of July, and on the northern coast of South America by August 21. Since the birds at Key West were fully 500 miles south of the breeding range, it is evident that they must have come from the southern part of the nesting area.
Figure 1.—Summer and winter homes of the black and white warbler, a very slow migrant as the birds nesting in the northern part of the country take 50 days to cross the breeding range. The speed of migration is shown in [figure 2]. ([See p. 14.])
Figure 2.—Isochronal migration lines of the black and white warbler, showing a very slow and uniform migration. These birds apparently advance only about 20 miles a day in crossing the United States. ([See p. 13.])
Many similar cases might be mentioned, such as the black-throated blue warblers, which are still observed in the mountains of Haiti in the middle of May, when others of this species are en route through North Carolina to breeding territory in New England or have even reached that region. Redstarts and yellow warblers, evidently the more southern breeders, are seen returning southward on the northern coast of South America just about the time that the earliest of those breeding in the North reach Florida on their way to winter quarters.