Segregation during migration
During the height of the northward movement in spring the woods and thickets may be suddenly filled with several species of wood warblers, thrushes, sparrows, flycatchers, and others, which it is natural to conclude have traveled together and arrived simultaneously. Probably they did, but such combined migration is by no means the rule for all species.
As a group the wood warblers (Compsothlypidae) probably travel more in mixed companies than do any other single family of North American birds. The flocks are likely to be made up of several species in spring and fall with both adults and young. Sometimes swallows, sparrows, blackbirds, and some of the shore birds also migrate in mixed flocks. In fall, great flocks of blackbirds frequently sweep south across the Plains States, and occasionally one flock will contain bronzed grackles, red-winged blackbirds, yellow-headed blackbirds, and Brewer's blackbirds.
On the other hand many species keep strictly to themselves. It would be difficult for any other kind of bird to keep in company with one of such rapid movements as the chimney swift, which is rarely found associated with any other species at any season. Nighthawks or bullbats also fly in separate companies, as do usually crows, waxwings, crossbills, bobolinks, and kingbirds. Occasionally, a flock of ducks will be observed to contain several species, but generally when they are actually on migration the individuals of each species separate and travel with others of their own kind. The flocks of blue geese, previously mentioned in connection with speed of flight (p. 22), frequently have with them a few of the closely related snow geese, particularly in the eastern part of their winter range. The portion here is usually about 10 to 1, but farther west the numbers of snow geese increase until they outnumber their blue relatives.
The adults of most perching birds drive the young away when they are grown, probably to be relieved of the necessity of providing for them, and also in order that the parents may have opportunity to rest and renew their plumage before starting for winter quarters. The young birds are therefore likely to drift together and, having no further responsibility, may start south ahead of their parents. In contrast with this indifference on the part of the adults of perching birds, Canada geese and some others remain in family groups, the parent birds undergoing the wing molt that renders them flightless during the period of growth of their young, so that old and young acquire their full plumage at the same time and are able to start south together. The large flocks, therefore, are composed of many families that band together, and when they separate into V-shaped units it is probably correct to assume that it is an old bird that leads the group. Where there is segregation of the sexes, the young birds usually accompany their mothers, as is the case with some of the ducks. After the females start to incubate their eggs, the males of most species of ducks flock by themselves and remain together until fall.
The males and females of some species may migrate either simultaneously or separately. In the latter case it is usually the males that arrive first, sometimes great flocks of male birds, as in the red-winged blackbird, reaching a locality several days before any of the females. This is particularly the rule in spring: The first robins are usually found to be males, as are also the first song sparrows, rose-breasted grosbeaks, and scarlet tanagers. This early arrival of the males has been explained on the theory of territorial possession, under which the male selects the area where it elects to breed, each individual attempting to protect a definite territory from trespass by other males of his own kind, at the same time singing or otherwise announcing his presence and inviting the later arriving female to examine the territory that he has selected for nesting. The long-billed marsh wren is a noteworthy example, and the males of this species may enthusiastically build several dummy nests before the females arrive.
In a few species the males and females apparently arrive at the breeding grounds together and proceed at once to nest building. In fact among the shore birds, ducks, and geese, courtship and mating may take place in whole or in part while the birds are in the South or on their way north, so that when they arrive at the northern nesting grounds they are paired and ready to proceed at once with the raising of their families. Mallards and black ducks may be observed in pairs as early as January, the female leading and the male following when they take flight. Naturally these mated pairs migrate north in company, and it was largely to protect such species that duck shooting in spring was abolished by Federal law a number of years ago.
Many shore birds nest well within the Arctic Circle, and it is the opinion of ornithologists that most of these birds share, at least in part, the habits of the phalaropes, a family in which the male assumes the entire care of the eggs and young. If this be true, it explains why in southern latitudes so many of the earliest fall arrivals are females that may have deserted the breeding grounds after the eggs were laid.
Migratory flights are frequently accomplished in close flock formation, as with the shore birds, blackbirds, and waxwings, and especially some of the sparrows—the snow buntings, longspurs, juncos, and tree sparrows. Other species, however, though they travel in flocks, maintain a very loose formation; examples are the turkey vultures, the hawks, swifts, blue jays, swallows, warblers, and bluebirds. Still others, the grebes, great horned owls, winter wrens, shrikes, and belted kingfishers for example, ordinarily travel alone and when several are found in close proximity it is an indication that they have been drawn together by unusual conditions, such as abundant food.
Where Birds Migrate
Definite evidence shows that both the length and the duration of the migratory journey vary greatly. The bobwhite and the western quails, the cardinal, the Carolina wren, and probably some of the titmice and woodpeckers, which are apparently almost or quite nonmigratory, may round out their full period of existence without at any time going more than 10 miles from the nest where they were hatched.