Migration of Birds (1950)

by Frederick Charles Lincoln

Wide and narrow migration lanes

When birds start their southward migration the movement necessarily involves the full width of the breeding range. Later there is a convergence of the lines of flight taken by individual birds, owing to the conformation of the land mass, and as the species proceeds southward the width of the occupied region becomes less and less. An example of this is provided by the common kingbird, which breeds from Newfoundland to British Columbia, a summer range 2,800 miles wide. On migration, however, its paths converge until in the southern part of the United States the occupied area extends from Florida to the mouth of the Rio Grande, a distance of only 900 miles, and still farther south the migration path is further restricted. In the latitude of Yucatan it is not more than 400 miles wide, and it is probable that the great bulk of the species moves in a belt that is less than half that width.

A migration route, therefore, may be anything from a narrow path that adheres closely to some definite geographical feature, such as a river valley or a coast line, to a broad boulevard that leads in the desired direction and which follows only the general trend of the land mass. Also it is to be remembered that whatever main routes are described, there remains a multitude of tributary and separate minor routes. In fact, with the entire continent of North America crossed by migratory birds, the different groups or species frequently follow lines that may repeatedly intersect those taken by others of their own kind or by other species. The arterial routes, therefore, must be considered merely as indicating paths of migration on which the tendency to concentrate is particularly noticeable.

In considering the width of migration lanes it will be obvious that certain species, as the knot and the purple sandpiper, which are normally found only along the coasts, must have extremely narrow routes of travel. They are limited on one side by the broad waters of the ocean, and on the other by land and fresh water, both of which are unsuited to furnish the food that is desired and necessary to the well-being of these species.

Figure 9.—Breeding and wintering ranges and migration of Harris's sparrow, an example of a narrow migration route through the interior of the country. The heavy broken lines enclose the region traversed by the majority of these finches; the light broken line encloses the country where they occur with more or less regularity; while the spots indicate records of accidental or sporadic occurrence.

Among land birds that have a definite migration, the Ipswich sparrow has what is probably the most restricted migration range of any species. It is known to breed only on Sable Island, Nova Scotia, and it winters along the Atlantic coast south to Georgia. Living constantly within sound of the surf, it is rarely more than a quarter of a mile from the outer beach, and is entirely at home among the sand dunes and their sparse covering of coarse grass.

Figure 10.—Distribution and migration of the scarlet tanager. During the breeding season individual scarlet tanagers may be 1,900 miles apart in an east-and-west line across the breeding range. In migration, however, the lines converge until in southern Central America they are not more than 100 miles apart. For migration paths of other widths see figures 9, 11, and 12.

Harris's sparrow supplies an interesting example of a narrow migration route in the interior of the country ([fig. 9.]) This fine, large finch is known to breed only in the region from Churchill, on the west shore of Hudson Bay, northwest to the shores of Great Bear Lake. Very few actual breeding records of the species are available, but these are sufficient to indicate that the breeding range is in the strip of country characterized by more or less stunted timber just south of the limit of trees. When it begins its fall migration, this species necessarily covers the full width of its breeding area. Then it proceeds almost directly south, or slightly southeasterly, the area covered by the majority of the birds becoming gradually constricted, so that by the time it reaches the United States it is most numerous in a belt about 500 miles wide, extending across North Dakota to central Minnesota. Harris's sparrows are noted on migration with fair regularity east to the western shore of Lake Michigan, and west to the foothills of the Rocky Mountains, but the great bulk of the species moves north and south through a relatively narrow path in the central part of the continent. Present knowledge suggests that the reason for this narrow migration range is the close association that Harris's sparrow maintains with a certain type of habitat' including brushy places, thickets, edges of groves, and weed patches. While these environmental conditions are found in other parts of the country, the region crossed by this sparrow presents almost a continuous succession of habitat of this type. Its winter range extends from southeastern Nebraska and northwestern Missouri, across eastern Kansas and Oklahoma and through a narrow section of central Texas, at places hardly more than 150 miles wide.

The scarlet tanager presents another extreme case of narrowness of migration route ([fig. 10]), its breeding range extending in greatest width from New Brunswick to Saskatchewan, a distance of about 1,900 miles. As the birds move southward in fall their path of migration becomes more and more constricted, until at the time they leave the United States all are included in the 600-mile belt from eastern Texas to the Florida peninsula. Continuing to converge through Honduras and Costa Rica, the boundaries there are not more than 100 miles apart. The species winters in northwestern South America, where it spreads out over most of Colombia, Ecuador, and Peru.

The rose-breasted grosbeak also leaves the United States through the 600-mile stretch from eastern Texas to Apalachicola Bay, but thereafter the lines do not further converge, as this grosbeak enters the northern part of its winter quarters in Central America and South America through a door of about the same width ([fig. 11]).

Although the cases cited represent extremes of convergence, a narrowing of the migratory path is the rule to a greater or less degree for the majority of North American birds. The shape of the continent tends to effect this, and so the width of the migration route in the latitude of the Gulf of Mexico is usually much less than in the breeding territory.

The redstart represents a notable case of a wide migration route, although even in the southern United States this is much narrower than the breeding range ([fig. 12]). These birds, however, cross all parts of the Gulf of Mexico and pass from Florida to Cuba and Haiti by way of the Bahamas, so that here their route has a width of about 2,500 miles.

Figure 11.—Distribution and migration of the rose-breasted grosbeak. Though the width of the breeding range is about 2,500 miles, the migratory lines converge until the boundaries are only about 700 miles apart when the birds leave the United States. For migration paths of other widths see figures 9, 10, and 12.