LIST OF ILLUSTRATIONS
- [A Fern Lover]
- [Prothallium Diagram]
- [Pinnate Frond]
- [Bipinnate Frond]
- [Pinnatifid Frond]
- [Spore Cases]
- [Linen Tester]
- [Curly Grass. Schizæa]
- [Cinnamon Fern. Osmunda cinnamomea]
- [Sensitive Fern. Onoclea sensibilis]
- [Ostrich Fern. Onoclea Struthiopteris]
- [Interrupted Fern. Osmunda Claytoniana]
- [Climbing Fern. Lygodium]
- [Flowering Fern. Osmunda regalis spectabilis]
- [Adder's Tongue. Ophioglossum]
- [Grape Fern. Botrychium]
- [Polypody. Polypodium]
- [Beech Fern. Phegopteris]
- [Cloak Fern. Notholæna]
- [Filmy Fern. Trichomanes]
- [Bracken. Pteris]
- [Maidenhair. Adiantum]
- [Cliff Brake. Pellæa]
- [Lip Fern. Cheilanthes]
- [Rock Brake. Cryptogramma]
- [Chain Fern. Woodwardia]
- [Shield Fern. Polystichum]
- [Wood Fern. Aspidium]
- [Bladder Fern. Cystopteris]
- [Woodsia]
- [Hayscented Fern. Dennstædtia]
- [Hart's Tongue. Scolopendrium]
- [Walking Fern. Camptosorus]
- [Asplenium Type]
- [Athyrium Type]
- [Sporangia of the Five Families]
- [Indusium]
- [Common Polypody. Polypodium vulgare]
- [Sori of Polypody]
- [Polypody in mass (Greenwood)]
- [Gray Polypody. Polypodium incanum]
- [Brake. Bracken. Sterile Frond]
- [Bracken. Fertile Frond]
- [Bracken, var. pseudocaudata]
- [Spray of Maidenhair]
- [Sori of Maidenhair]
- [Maidenhair. Adiantum pedatum]
- [Alpine Maidenhair]
- [Venus-Hair Fern. Adiantum capillus-veneris]
- [Purple Cliff Brake. Pellæa atropurpurea]
- [Dense Cliff Brake. Cryptogramma densa]
- [Slender Cliff Brake. Cryptogramma Stelleri]
- [Parsley Fern. Cryptogramma acrostichoides]
- [Alabama Lip Fern. Cheilanthes alabamensis]
- [Hairy Lip Fern. Cheilanthes lanosa]
- [Slender Lip Fern. Cheilanthes Féei]
- [Pinnæ of Slender Lip Fern]
- [Powdery Cloak Fern. Notholæna dealbata]
- [Common Chain Fern. Woodwardia virginica]
- [Net-veined Chain Fern. Woodwardia areolata]
- [The Spleenworts]
- [Pinnatifid Spleenwort. Asplenium pinnatifidum]
- [Scott's Spleenwort. Asplenium ebenoides]
- [Green Spleenwort. Asplenium viride]
- [Maidenhair Spleenwort. Asplenium Trichomanes]
- [Maidenhair Spleenwort. Asplenium Trichomanes (Fernery)]
- [Ebony Spleenwort. Asplenium platyneuron]
- [Bradley's Spleenwort. Asplenium Bradleyi]
- [Mountain Spleenwort. Asplenium montanum]
- [Rue Spleenwort. Asplenium Ruta-muraria]
- [Rootstock of Lady Fern (Two parts)]
- [Sori of Lady Fern. Athyrium angustum]
- [Varieties of Lady Fern]
- [Lowland Lady Fern. Athyrium asplenioides]
- [Silvery Spleenwort. Athyrium acrostichoides]
- [Narrow-leaved Spleenwort. Athyrium angustifolium]
- [Pinnæ and Sori of Athyrium angustifolium]
- [Sori of Scolopendrium vulgare]
- [Hart's Tongue. Scolopendrium vulgare]
- [Walking Fern. Camptosorus rhizophyllus]
- [Christmas Fern. Polystichum acrostichoides]
- [Varieties of Christmas Fern]
- [Braun's Holly Fern. Polystichum Braunii]
- [Holly Fern. Polystichum Lonchitis]
- [Marsh Fern. Aspidium Thelypteris]
- [Marsh Fern, in the mass]
- [Massachusetts Fern. Aspidium simulatum]
- [New York Fern. Aspidium noveboracense]
- [Sori of Aspidium noveboracense]
- [Pinnæ and Sori of Aspidium noveboracense]
- [Oak Fern. Phegopteris Dryopteris]
- [Northern Oak Fern. Phegopteris Robertiana]
- [Broad Beech Fern. Aspidium hexagonoptera]
- [Long Beech Fern. Aspidium polypedioides]
- [Fragrant Fern. Aspidium fragrans]
- [Marginal Shield Fern. Aspidium marginale]
- [Crown of Fronds of Aspidium marginale]
- [Sori of Aspidium marginale]
- [Male Fern. Aspidium Filix-mas]
- [Aspidium Filix-mas and details]
- [Goldie's Shield Fern. Aspidium Goldianum]
- [Aspidium Goldianum, in the mass]
- [Crested Shield Fern. Aspidium cristatum]
- [Crested Shield Fern. Aspidium cristatum (No. 2)]
- [Clinton's Shield Fern. Aspidium cristatum var. Clintonianum]
- [Crested Marginal Fern. Aspidium cristatum × marginale]
- [Aspidium cristatum × marginale, in the mass]
- [Boott's Shield Fern. Aspidium Boottii]
- [Spinulose Shield Fern. Aspidium spinulosum]
- [Aspidium spinulosum var. intermedium]
- [Aspidium spinulosum var. americanum]
- [Bulblet Bladder Fern. Cystopteris bulbifera]
- [Cystopteris bulbifera with sprouting bulb]
- [Fragile Bladder Fern. Cystopteris fragilis]
- [Rusty Woodsia. Woodsia ilvensis]
- [Northern Woodsia. Woodsia alpina]
- [Details of Alpine Woodsia]
- [Blunt-lobed Woodsia. Woodsia obtusa]
- [Smooth Woodsia. Woodsia glabella]
- [Hayscented Fern. Dennstædtia punctilobula]
- [Forked variety of Dennstædtia punctilobula]
- [Field View of Dennstædtia punctilobula]
- [Pinnæ and Sori of Dennstædtia punctilobula]
- [Meadow View of Sensitive Fern]
- [Obtusilobata Forms of Sensitive Fern, Leaf to Fruit]
- [Sori of Sensitive Fern]
- [Sensitive Fern. Onoclea sensibilis]
- [Sensitive Fern, Fertile and Sterile Fronds on Same Plant]
- [Ostrich Fern. Onoclea Struthiopteris. Fertile Fronds]
- [Ostrich Fern. Sterile Fronds]
- [Sori and Sporangia of Ostrich Fern]
- [Royal Fern. Osmunda regalis spectabilis]
- [Sori of Royal Fern]
- [Interrupted Fern. Osmunda Claytoniana]
- [Interrupted Fern. Fertile Pinnules Spread Open]
- [Cinnamon Fern. Osmunda cinnamomea]
- [Cinnamon Fern. Leaf Gradations]
- [Two Varieties of Cinnamon Fern]
- [Osmunda cinnamomea glandulosa]
- [Curly Grass. Schizæa pusilla]
- [Sporangia of Curly Grass]
- [Climbing Fern. Lygodium palmatum]
- [Adder's Tongue. Ophioglossum vulgatum]
- [Moonwort. Botrychium Lunaria]
- [Moonwort, Details]
- [Little Grape Fern. Botrychium simplex]
- [Lance-leaved Grape Fern. Botrychium lanceolatum]
- [Matricary Grape Fern. Botrychium ramosum]
- [Common Grape Fern. Botrychium obliquum]
- [Botrychium obliquum var. dissectum]
- [Botrychium obliquum var. oneidense]
- [Ternate Grape Fern. Botrychium ternatum var. intermedium]
- [Ternate Grape Fern. B. ternatum var. intermedium]
- [Rattlesnake Fern. Botrychium virginianum]
- [Filmy Fern. Trichomanes Boschianum]
- [Fruiting Pinnules of Filmy Fern]
- [Crosiers]
- [Noted Fern Authors]
- [Spray of the Bulblet Bladder Fern]
[PREFACE]
A lover of nature feels the fascination of the ferns though he may know little of their names and habits. Beholding them in their native haunts, adorning the rugged cliffs, gracefully fringing the water-courses, or waving their stately fronds on the borders of woodlands, he feels their call to a closer acquaintance. Happy would he be to receive instruction from a living teacher: His next preference would be the companionship of a good fern book. Such a help we aim to give him in this manual. If he will con it diligently, consulting its glossary for the meaning of terms while he quickens his powers of observation by studying real specimens, he may hope to learn the names and chief qualities of our most common ferns in a single season.
Our most productive period in fern literature was between 1878, when Williamson published his "Ferns of Kentucky," and 1905, when Clute issued, "Our Ferns in Their Haunts." Between these flourished D.C. Eaton, Davenport, Waters, Dodge, Parsons, Eastman, Underwood, A.A. Eaton, Slosson, and others. All their works are now out of print except Clute's just mentioned and Mrs. Parsons' "How to Know the Ferns." Both of these are valuable handbooks and amply illustrated. Clute's is larger, more scholarly, and more inclusive of rare species, with an illustrated key to the genera; while Mrs. Parsons' is more simple and popular, with a naive charm that creates for it a constant demand.
We trust there is room also for this unpretentious, but progressive, handbook, designed to stimulate interest in the ferns and to aid the average student in learning their names and meaning. Its geographical limits include the northeastern states and Canada. Its nomenclature follows in the main the seventh edition of Gray's Manual, while the emendations set forth in Rhodora, of October, 1919, and also a few terms of later adoption are embodied, either as synonyms or substitutes for the more familiar Latin names of the Manual, and are indicated by a different type. In every case the student has before him both the older and the more recent terms from which to choose. However, since the book is written primarily for lovers of Nature, many of whom are unfamiliar with scientific terms, the common English names are everywhere given prominence, and strange to say are less subject to change and controversy than the Latin. There is no doubt what species is meant when one speaks of the Christmas fern, the ostrich fern, the long beech fern, the interrupted fern, etc. The use of the common names will lead to the knowledge and enjoyment of the scientific terms.
A friend unfamiliar with Latin has asked for pointers to aid in pronouncing the scientific names of ferns. Following Gray, Wood, and others we have marked each accented syllable with either the grave (`) or acute (´) accent, the former showing that the vowel over which it stands has its long sound, while the latter indicates the short or modified sound. Let it be remembered that any syllable with either of these marks over it is the accented syllable, whose sound will be long or short according to the slant of the mark.
We have appropriated from many sources such material as suited our purpose. Our interest in ferns dates back to our college days at Amherst, when we collected our first specimens in a rough, bushy swamp in Hadley. We found here a fine colony of the climbing fern (Lygodium). We recall the slender fronds climbing over the low bushes, unique twiners, charming, indeed, in their native habitat. We have since collected and studied specimens of nearly every New England fern, and have carefully examined most of the other species mentioned in this book. By courtesy of the librarian, Mr. William P. Rich, we have made large use of the famous Davenport herbarium in the Massachusetts Horticultural library, and through the kindness of the daughter, Miss Mary E. Davenport, we have freely consulted the larger unmounted collection of ferns at the Davenport homestead, at Medford,[1] finding here a very large and fine assortment of Botrychiums, including a real B. ternatum from Japan.
[Footnote 1: Recently donated to the Gray Herbarium.]
For numerous facts and suggestions we are indebted to the twenty volumes of the Fern Bulletin, and also to its able editor, Mr. Willard N. Clute. To him we are greatly obligated for the use of photographs and plates, and especially for helpful counsel on many items. We appreciate the helpfulness of the American Fern Journal and its obliging editor, Mr. E.J. Winslow. To our friend, Mr. C.H. Knowlton, our thanks are due for the revision of the checklist and for much helpful advice, and we are grateful to Mr. S.N.F. Sanford, of the Boston Society of Natural History, for numerous courtesies; but more especially to Mr. C.A. Weatherby for his expert and helpful inspection of the entire manuscript.
The illustrations have been carefully selected; many of them from original negatives bequeathed to the author by his friend, Henry Lincoln Clapp, pioneer and chief promoter of school gardens in America. Some have been photographed from the author's herbarium, and from living ferns. A few are from the choice herbarium of Mr. George E. Davenport, and also a few reprints have been made from fern books, for which due credit is given. The Scott's spleenwort, on the dedication page, is reprinted from Clute's "Our Ferns in Their Haunts."
[INTRODUCTION]
Thoreau tells us, "Nature made a fern for pure leaves." Fern leaves are in the highest order of cryptogams. Like those of flowering plants they are reinforced by woody fibres running through their stems, keeping them erect while permitting graceful curves. Their exquisite symmetry of form, their frequent finely cut borders, and their rich shades of green combine to make them objects of rare beauty; while their unique vernation and method of fruiting along with their wonderful mystery of reproduction invest them with marked scientific interest affording stimulus and culture to the thoughtful mind. By peculiar enchantments these charming plants allure the ardent Nature-lover to observe their haunts and habits.
"Oh, then most gracefully they wave
In the forest, like a sea,
And dear as they are beautiful
Are these fern leaves to me."
As a rule the larger and coarser ferns grow in moist, shady situations, as swamps, ravines, and damp woods; while the smaller ones are more apt to be found along mountain ranges in some dry and even exposed locality. A tiny crevice in some high cliff is not infrequently chosen by these fascinating little plants, which protect themselves from drought by assuming a mantle of light wool, or of hair and chaff, with, perhaps, a covering of white powder as in some cloak ferns--thus keeping a layer of moist air next to the surface of the leaf, and checking transpiration.
Some of the rock-loving ferns in dry places are known as "resurrection" ferns, reviving after their leaves have turned sere and brown. A touch of rain, and lo! they are green and flourishing.
Ferns vary in height from the diminutive filmy fern of less than an inch to the vast tree ferns of the tropics, reaching a height of sixty feet or more.