IV.
It was, as we have said, in 1868 that Darwin published the two volumes collectively entitled "Variation of Animals and Plants under Domestication." It is the second and largely corrected edition brought out in 1875 which we have under our eye. It is the outcome of the views maintained by the author in this work and elsewhere that not only the various domestic races but the most distinct genera and orders within the same great class--for instance, mammals, birds, reptiles, and fishes--are all the descendants of one common progenitor, and the whole vast amount of difference between these forms has primarily arisen from simple variability. Darwin recognized that he who for the first time should consider the subject under this point of view would be struck dumb with amazement. He submits, however, that the amazement ought to be lessened when we reflect that beings almost infinite in number during an almost infinite lapse of time have often had their whole organization rendered in some degree plastic, and that each slight modification of structure which was in any way beneficial under excessively complex conditions of life has been preserved, whilst each which was in any way injurious has been rigorously destroyed. The long-continued accumulation of beneficial variations will infallibly have led to structures as diversified, as beautifully adapted for various purposes, and as excellently co-ordinated as we see in the animals and plants around us. Hence Darwin regards selection as the paramount power, whether applied by man to the formation of domestic beings or by nature to the production of species. Employing a favorite metaphor, he said: "If an architect were to rear a noble and commodious edifice without the use of cut stone, by selecting from the fragments at the base of a precipice wedge-form stones for his arches, elongated stones for his lintels, and flat stones for his roof, we should admire his skill and regard him as the paramount power. Now, the fragments of stone, though indispensable to the architect, bear to the edifice built by him the same relation which the fluctuating variations of organic beings bear to the varied and admirable structures ultimately acquired by their modified descendants."
Some critics of the Darwinian theory of the origin of species have declared that natural selection explains nothing, unless the precise cause of each slight individual difference be made clear. Darwin rejoins that if it were explained to a savage utterly ignorant of the art of building how the edifice had been raised, stone upon stone, and why wedge-formed fragments were used for the arches, flat stones for the roof, etc.; and if the use of each part and of the whole building were pointed out,--it would be unreasonable if he declared that nothing had been made clear to him, because the precise cause of the shape of each fragment could not be told. This, in Darwin's opinion, is a nearly parallel case, with the objection that selection explains nothing because we know not the cause of each individual difference in the structure of each being. The shape of the fragments of stone at the base of the hypothetical precipice may be called accidental, but the term is not strictly applicable; for the shape of each depends on a long sequence of events, all obeying natural laws; on the nature of the rock, on the lines of deposition or cleavage, on the form of the mountain, which depends on its upheaval and subsequent denudation, and, lastly, on the storm or earthquake which throws down the fragments.
In regard to the use, however, to which the fragments may be put, their shape may be strictly said to be accidental. Here Darwin acknowledged that we are brought face to face with a great difficulty in alluding to which he felt that he was travelling beyond his proper province. "An omniscient Creator must have foreseen every consequence which results from the laws imposed by Him. But can it be reasonably maintained that the Creator intentionally ordered, if we use the words in any ordinary sense, that certain fragments of rock should assume certain shapes, so that the builder might erect his edifice? If the various laws which have determined the shape of each fragment were not predetermined for the builder's sake, can it be maintained with any greater probability that He specially ordained for the sake of the breeder each of the innumerable variations in our domestic animals and plants,--many of these variations being of no service to man, and not beneficial, far more often injurious, to the creatures themselves? Did He ordain that the crop and tail-feathers of the pigeon should vary in order that the fancier might make his grotesque pouter and fan-tail breeds? Did He cause the frame and mental qualities of the dog to vary in order that a breed might be formed of indomitable ferocity with jaws fitted to pin down the bull for man's brutal sport?"
It is obvious, however, that if we give up the principle in one case,--if we do not admit that the variations of the primeval dog were intentionally guided in order that the greyhound, for instance, that perfect image of symmetry and vigor, might be formed,--no shadow of reason can be assigned for the belief that variations similar in nature and the result of the same general laws which have been the groundwork through natural selection of the formation of the most perfectly adapted animals in the world, man included, were intentionally and specially guided. Darwin, therefore, was unable to follow the distinguished botanist, Prof. Asa Gray, in his belief that "variation has been led along certain beneficial lines," like a stream "along definite and useful lines of irrigation." Darwin's conclusion was that, if we assume that each particular variation was from the beginning of all time preordained, then that plasticity of organization which leads to many injurious deviations of structure, as well as the redundant power of reproduction which inevitably leads to a struggle for existence, and, as a consequence, to a natural selection or survival of the fittest, must appear to us superfluous laws of nature.