DIGESTION
Definition. Digestion is the breaking up, changing, and liquefying of the food in the various chambers of the alimentary canal designed for that purpose. The mechanical breaking up is done principally by the teeth in the mouth, the chemical changes and liquefying by the various digestive fluids.[19]
Digestive Fluids. The digestive fluids are true secretions. Each is formed from the blood by a special gland for the purpose which never does anything else; they do not exist in the blood as such. Their flow is intermittent, taking place only when they are needed. The liver, however, is an exception to all the others. It is both secretory and excretory, and bile is formed all the time, but is most abundant during digestion.[20]
Saliva. The fluid which is mixed with the food in the mouth is secreted by a considerable number and variety of glands, the principal of which are the parotid, submaxillary, and sublingual. Smaller glands in the roof and sides of the mouth, in the tongue, and in the mucous membrane of the pharynx contribute to the production of saliva, the digestive fluid of the mouth. The flow from the parotid gland is greatest. The flow from all the glands is greatly increased when food is taken, especially if it be of good flavor. Sometimes the amount is increased by smell alone, as when a nice steak is cooking, or a savory soup, and sometimes the saliva is made copious by thought, as when we remember the taste of dishes eaten in the past, and we say, "It makes the mouth water just to think of them."
Amount of Saliva. According to Dalton the amount of saliva secreted every twenty-four hours is 42½ oz. Its reaction is almost constantly alkaline. It is composed of water, organic matter, and various mineral salts. Ptyalin is its active principle, and is called by some authors animal diastase, or starch converter.
Gastric Juice. Gastric juice is the digestive fluid of the stomach. It is acid. Its flow is intermittent, occurring only at times of digestion. Its active principle is pepsin.
It is worthy of notice here that the character of the digestive fluids when food is taken is different from what it is when the organs are at rest. For instance, the gastric juice which flows in abundance under the stimulus of food, is not like the fluid secreted when the stomach is collapsed and empty.
Pancreatic Juice. Pancreatic juice is the digestive juice of the pancreas, and is poured into the small intestine a short distance below the pyloric opening. Its reaction is alkaline. Its flow is entirely suspended during the intervals of digestion.
Bile. Bile, the fourth in order of the digestive liquids, is the secretion of the largest gland of the body—the liver. It is poured into the small intestine by a duct which empties side by side with the duct from the pancreas. The flow of bile is constant, but is greatest during digestion.
Intestinal Juice. Intestinal juice has been to physiologists a difficult subject of study. It is mingled with the salivary and gastric juices at the times of digestion, when it is most desirable to notice its action. Nearly all authorities agree that it is alkaline, and that its function is to complete the digestion of substances which may reach it in an undigested condition.
Mucus of Large Intestine. The mucus secreted by the large intestine is for lubricating only.
Digestion in Different Parts of the Alimentary Tract. Different substances in food are digested in different portions of the alimentary canal, and by different means. Let us begin in the mouth. Taking the classes of foods, starch, one of the carbohydrates, is the one most affected by the ptyalin, or animal diastase, of the saliva. So energetic is the action of ptyalin on starch that 1 part is sufficient to change 1000 parts. Starch is not acted upon by the gastric juice of the stomach at all; however, the continued action of the saliva is not probably interrupted in the stomach. The digestion of starch is completed by the action of the pancreatic and intestinal juices, and consists in its being changed into soluble glucose, which is absorbed in solution.
Sugar. Cane-sugar, or common sugar (also called sucrose), passes through the mouth, unchanged, to the stomach, where it is converted into glucose by the slow action of the acid (hydrochloric) of the gastric juice. Dilute hydrochloric acid has the same action on sugar outside of the stomach.
The action of pancreatic juice on sugar is very marked; it immediately changes cane-sugar into glucose. The effect of intestinal fluid is not well understood, but there is the general agreement that it does not change cane-sugar, neither is cane-sugar, as such, absorbed in the intestine. Bile does not affect it, therefore cane-sugar is digested or converted into glucose either by the stomach or pancreas, or both. It will now be seen that ultimately the same substance, glucose, is obtained from both starch and sugar.
Protein. We now come to the consideration of the digestion of the protein compounds, of which albumen may be taken as a type. Possibly no action except breaking up and moistening takes place in the mouth.[21] Its digestion begins in the stomach, where its structure is broken up and a separation and dissolution of the little sacs which hold it take place. The same thing is partially accomplished outside of the stomach when white of egg is slightly beaten and strained through a cloth. Gastric juice further acts on the albumen itself, forming it into what is called albumen peptone. The digestion of raw and carefully cooked albumen has been found to be carried on very rapidly in the stomach, and the change is essentially the same in both cases, but in favor of the slightly coagulated. When the albumen is rendered hard, fine, and close in consistency by over-cooking, then it is less easy of digestion than when raw.
Absorption. It is probable that the greater portion of the process of digestion and absorption of albumen takes place in the stomach.
Fibrin. Fibrin is also digested in the stomach, and made into fibrin peptone.
Casein. Liquid casein is immediately coagulated by gastric juice, both by the action of free acid and organic matter.
Gelatin. Gelatin is quickly dissolved by gastric juice, and afterward no longer has the property of forming jelly on cooling. Gelatin is more rapidly disposed of than the tissue from which it is produced.
Vegetable Protein. The digestion of the vegetable protein compounds, such as the gluten of wheat and the protein of the various grains, such as corn, oatmeal, etc., is undoubtedly carried on in the stomach, but they must be well softened and prepared by the action of heat and water, or they will not be digested anywhere; and often corn, beans, and grains of oatmeal are rejected entirely unchanged. Partially or imperfectly digested proteins are affected by intestinal juice. It is probable that the function of this fluid is to complete digestive changes in food which have already begun in the stomach.
To summarize: The digestion and absorption of nitrogenous compounds take place in both the stomach and the intestines.