Botany: The Science of Plant Life

by Norman Taylor

2. How Plants Get Their Food and Water From the Earth

If we could stretch an apparently impervious membrane, like the inner white skin just inside an eggshell, or a piece of parchment, and so form a wall through the middle of a glass box, and then pour into one of the compartments pure water and in the other a mixture of water and molasses, a very curious result would follow within a comparatively short period. We should find that presently there would be a gentle filtering of the water through the membrane toward the molasses water, and similar gentle current in the other direction. In other words, fluids of different density, if separated by a membrane, tend to equalize each other. This equalization may not be very rapid, and at first it will be more speedy from the less dense to the more dense, but eventually it will make the different fluids of a common density. This purely mechanical property of the equalization of fluids separated by a membrane is known as osmosis, and it is upon the possession of the equipment necessary for this that roots depend for getting food and water from the soil.

In our discussion of roots in Chapter I, we found that they end in very fine subdivisions, which are themselves split up into practically invisible root hairs. These root hairs are the only way that roots can absorb the food and water in the soil, and they are able to do this because they are provided with a membrane which permits osmosis to act between the solution inside the root hair and the water in the soil. The solution in the root hair is mostly a sugary liquid, some of that surplus sugar made in the leaves, and it is denser than the soil water, so there is apparently nothing to prevent an equalization of the liquids on different sides of the membrane. If this actually happened, as it would in the case of the simple experiment noted above, then roots would exchange a fairly rich sugary liquid for a much more watery one, and we should find that plants did not get their food from the soil, but really have it drained away from them by osmosis. But nature has a cunning device for stopping such robbery, which is prevented by the membranes of root hairs being only permeable to the extent of letting water in, not permeable enough to allow sugar to escape. As we have seen, osmosis is a purely mechanical process which, if left to operate without interference, would not aid but injure the plant. Surely, nothing with which plants are provided is so important to them as this delicate membrane of the root hairs which, while allowing osmosis to act in a one-sided fashion, preserves to the plant the sugary liquid that alone makes the absorption of soil water possible.

As root hairs are very much alive and work constantly, they must be provided with air, without which no living thing can exist. And here, again, it seems as though nature, with almost uncanny foresight, had deliberately planned for this requirement of roots. And, in this case, not by interfering with a physical process by an adjustment of plant structure, but by the arrangement of soil particles and the way in which water is found in all soils. Soil particles, even in the most compact clay, do not fill all the space occupied by the soil as a whole. There are tiny air spaces all through the soil, which insures a constant supply of fresh air. That is one reason why gardens are cultivated, to see to it that plenty of fresh air is allowed to permeate the soil. Around the finest soil particles there is always an almost incredibly thin film of water, which is renewed as soon as it is lost by its absorption by the root hairs or by evaporation. This renewal of the water film is itself a mechanical process, called capillarity, best illustrated by putting a few drops of water on a plate and placing on them a lump of sugar. The water will spread all through the lump of sugar in a few seconds and the capillarity that forces it up through the lump is the same as sees to it that the tiny film of water surrounding the finest soil particles is constantly renewed from the lower levels of the soil.

Little do we dream, as we walk over the commonest weed, that buried at its roots are these delicate arrangements for securing food and water. Osmosis allowed to act so that the “exchange” of liquids is all to the advantage of the plant, capillarity providing a constant water supply, and the very piling together of the soil so contrived that the life-giving air filters all through it—does it not seem as if all this were, if not a deliberate plan, certainly a more perfect one than mere man could have devised?

If you will turn back for a moment to the beginning of the description of how plants get their food, you will find that in osmosis the weaker liquid tends to permeate the denser one more rapidly than the denser one does the weaker. As we have just seen, the sugary liquid in the root hairs is denser than the soil water outside, and, furthermore, none of it is allowed to escape. This comparatively greedy process of taking everything and giving nothing results in a constant flow of soil water into the root hairs. When the flow of liquids in osmosis is not at once equalized, a gentle pressure is brought to bear to make them so. This is what is called osmotic pressure, and it is this pressure that forces the absorbed liquid through the roots and part way up the trunk of even the tallest trees. While we have just said it is a gentle pressure, that is true only in the case where the osmosis has free play, and the pressure is stopped with the perfect mixing of the two liquids. But what if they can never mix? What may not the accumulated osmotic pressure amount to in such a one-sided process as goes on in root hairs with everything coming in and nothing going out. Cut-off stems, with a pressure gauge attached to them, indicate that in some plants the pressure is from 60 up to 170 pounds!

Another result of this pressure is that it keeps leaves and the fleshy stems of plants in their ordinary position. The actual solid part of nearly all leaves is scarcely 5 per cent of their bulk and all the rest is water. The constant pressure of this water from the roots is sufficient to keep leaves comparatively stiff and rigid, how stiff is quickly realized if the pressure stops and the leaf wilts or withers. Sometimes this osmotic pressure, particularly during rainy weather, becomes so great as to cause injury to the plant, the splitting of tomatoes and occasionally of plums, being due to it. This osmotic pressure, together with the extra pull given by the leaves, is sufficient to account for the rise of water to the tops of the tallest trees. The tallest trees in the world are certain kinds of blue gum in Australia which frequently reach a height exceeding 300 feet. What the combined osmotic pressure and leaf pull must be to carry such a heavy thing as water to such a great height is easier to imagine than to calculate.

The root hairs, then, by the process already described, absorb the water from the soil, but plants can no more live on water alone than we can. As we have seen, the membrane in the root hairs cannot allow the passage of even the tiniest particle of solid matter. In fact the root hair itself is so small that it can only be seen through the microscope, and of course the membrane is smaller still. Plant foods, then, can never be solids, but must always be such materials as can be dissolved in water. The chief of these are chemical substances, such as lime, potash, nitrogen, magnesium, phosphorus, sulphur, and iron. Hydrogen is also necessary, but as this makes up half the composition of water, there is a permanent supply of that provided by the soil water. These things make up the great part of plant foods taken in through the roots, and it is from these that leaves, by a process you already understand in its essential details, manufacture sugar and starch.

But neither starch nor sugar, important as they are to the plant, and absolutely necessary as they are to us, are the only things made by plants. Leaves may well be called factories, but plants are themselves the most wonderful chemical laboratories, beside which any built by man are as play-things. For plants, by processes too complicated to be explained here, work over their accumulation of starch and sugar, recombine some of their constituents, and store up in various parts of the plant the results, which are often such food ingredients as protein. This is the really essential food substance in wheat, as it is in eggs and meat. No chemist has ever succeeded in making a single scrap of it, yet it is such an everyday occurrence in practically all plants that it, with starch and sugar, forms the great food supply of the world. Not protein alone, but all the amazing plant products like the oils from the olive and the resin from pine, rubber, the drugs of plant origin, even tobacco—all these and hundreds of others are made by plants from those few simple foods absorbed through the roots, literally pumped up to the leaves and there, under the magic of sunlight, combined and recombined, worked over and changed utterly in their make-up. Nothing could be more perfect than the marshaling of forces and contrivances to secure the result; let there be even the least bungling, and for us the world would cease to be worth fighting for.

Nor does the work of plants stop here. If it did, they would be not unlike a commission merchant who had gathered from the four corners of the earth a supply of eggs only to find he could not or more likely would not sell them all at once, and yet had failed to provide himself with proper storage. Plants, too, have times in their life when adequate storage is necessary for them. So true is this that unless there is food enough stored in seeds to give a start to seedlings before their own roots begin to work, they would die almost at once. In seeds and in many nearly dormant parts of plants these foods are stored away for future use. The tubers of potatoes and all our root crops, like beet and parsnip, are common examples of this. Even the manufacture of wood in the trunks of trees is a storage appliance on the part of the plant, for wood is just as much one of the food products of a plant as wheat or rice.