PLANT FAMILIES AND ORDERS
A scientist once visiting in Bulgaria noticed that the peasants in that country frequently lived over a hundred years and, in trying to find out the reason, he discovered that they drank large quantities of sour milk. This is alive with a definite kind of bacterium that is of great benefit to the digestive apparatus, and therefore helps in the prolongation of life. In Bulgaria, in other words, a certain food habit of the people has resulted in a definite prolongation of life and fixes that population as of somewhat different characteristics from people not addicted to sour milk. In Japan a whole race lives largely on fish and rice, and while this is not the cause of their yellow skin, it is almost surely the cause of their generally small stature. Many of the English are tall, light-haired, and blue-eyed people, fond of outdoor life and sports, and among the most highly developed of the peoples of the earth. The climate of that island, their generally large consumption of meat and the outdoor life of so many of them, have resulted in quite definite characteristics that make the typical Englishman an easily distinguishable type.
In studying man we are able not only to divide him into such broad divisions as white, black, and yellow races, but due to their particular country or mode of life there are scores of racial subdivisions of these larger categories that everyone recognizes. Such differences are often based on stature, shape of head, mental characters and many others, but those still finer shades of difference between, for instance, a Connecticut Yankee and a plantation owner in the South, are, while noted by everyone, very difficult to accurately describe.
In attempting to find such major differences in plants, some structural character that would set off one large group of plants from every other group, the botanist has a harder task than the person studying man. For all those differences of language and mentality that make up such a large part of our common knowledge of the different peoples of the earth are characters that are foreign to plants. We are thus thrown back on structure as the chief way in which plants differ, and because their reproductive organs are their most important ones, and therefore least likely to vary, it is upon certain characters of these organs that all flowering plants have been divided.
In the chapter on “How Plants Produce Their Young,” we found that most flowering plants have their ovules in an ovary which, after fertilization, develop into fruit and seed. But some plants, while they have ovules, only bear them naked or between scales, never inclosed in an ovary. This is true in all pines, spruces, hemlocks, and all the host of their generally evergreen relatives. Such trees bear cones, between the scales of which are perfectly naked ovules that develop into seeds ([Figure 77]) that have never been hidden in an ovary, as have the vast majority of the seeds of other plants ([Figure 53]). These naked-seeded plants are known as gymnosperms or literally gymnos, naked, and sperma, seed, and comprise all the cone-bearing trees in the world, the larger part of which are always evergreen. In some past ages such trees made up the bulk of vegetation of the earth, but at present they are much reduced in numbers. Familiar examples of these Coniferæ, or cone-bearing trees, are larch, spruce, fir, pine, hemlock, juniper, and yew.
Most of these are evergreen, which does not, of course, mean that they bear the same leaves always, but that only a few drop off at a time and are so constantly renewed that the tree is actually ever green.
All other flowering plants always bear their ovules in an ovary and, because of this fact, are called angiosperms, literally angeion, a vessel, and sperma, seed. These inclosed seeded plants comprise the great bulk of the vegetation of the earth to-day. So far as the temperate zone is concerned, nearly all of them drop their leaves in the fall, and the trees belonging to the angiosperms are thus said to be deciduous trees.
No better idea of the present size and importance of these two groups of plants can be gained than to state the fact that perhaps not over 500 different kinds of gymnosperms, all of which are trees and shrubs, are known. All the rest of the flowering plants in the world, comprising over 150,000 different kinds of herbs, shrubs, and trees of infinite variety, are angiosperms and therefore bear ovules in an ovary, followed by seeds in or on some sort of a fruit. It would almost seem as though the simplest way to dispose of this great mass of plants would be to sort them into trees, shrubs, and herbs. For all of them belong to one of these types of plant growth, and the ancient students of plants, just before the time of Christ, actually divided all flowering plants into these three classes. This, of course, threw the coniferous trees in with all other kinds and, as we have already seen, they differ from all other kinds in the important character of having naked ovules.
Here, again, in order to get some system out of apparent chaos, we must fall back on some fundamental character. And, again, it is the product of the reproductive process in all this host of angiosperms which furnishes the clue. In the seeds of many of them the young embryo has folded up within it two seed leaves, while in all the rest only one. As we saw in Chapter I, these seeds germinate either with a single seed leaf, like corn ([Figure 85]), or with two seed leaves, like beans ([Figure 81]). Every one of these angiosperms belongs to one of these classes or the other, and perhaps more extraordinary still is the fact that in those with one seed leaf there are associated certain leaf and flower characters, while those with two seed leaves are always very different.
In the monocotyledons, or plants with a single seed leaf, the leaves are practically always parallel veined ([Figure 83]), like corn and grass, and lilies and palms, and hundreds of others. Also, they nearly always have the parts of their flowers in threes ([Figure 84]). That is, they have three sepals, petals, stamens, and often pistils, or multiples of three. The common trillium or wake-robin, for instance, has three sepals, three petals, six stamens, and three styles. With a few exceptions, and nature seems to delight in producing a few such, all monocotyledons have this parallel-veined leaf character and flower parts in threes or multiples of three.
Plants which send up two seed leaves ([Figure 81]), on the other hand, bear practically always netted-veined leaves ([Figure 79]), and the parts of their flowers are nearly always in fours or fives or multiples of these numbers ([Figure 80]). The well-known wild geranium has five sepals, five petals, ten stamens, and a five-lobed or five-celled ovary. There is some individual variation from this plan, sometimes one organ and sometimes another having more or less than the regular number. But so overwhelmingly true are these distinctions that dicotyledons, or plants with two seed leaves, and monocotyledons,
Dicotyledonous and Monocotyledonous growth habits contrasted. Figs. 78-81. The trunk of a dicotyledonous tree showing division of the wood into heartwood, sapwood, and cambium, which the removal of a piece of outer bark exposes. Note the net-veined leaf (79), the seedling with two seed leaves (81), and with the parts of the flower in 5’s (80). Figs. 82-85. Monocotyledonous plant. Note the lack of zones of wood, cambium and corky bark. Such plants have parallel-veined leaves (83), parts of their flowers in 3’s or 6’s (84), and germinate with a single seed leaf (85).
or plants with a single seed leaf, have been for hundreds of years the two great classes into which all angiospermous flowering plants are divided.
Our general view of all the flowering plants may be summarized then as follows:
1. Gymnosperms, or naked seeded plants, include all cone-bearing plants, mostly evergreen and always trees or shrubs. The pine is a familiar example.
2. Angiosperms, or inclosed seeded plants, include all other flowering plants of whatever kind. Divided into: (a) Monocotyledons. Sprouting with one seed leaf, and leaves practically always parallel-veined. Parts of the flower in threes or multiples of three. Familiar examples are corn, grass, sugar-cane, palms, cannas, and lily of the valley ([Figures 82-85]). (b) Dicotyledons. Sprouting with two seed leaves, and the leaves practically always netted-veined. Parts of the flower in fours or fives or multiples of these numbers. Includes all the remaining flowering plants and is a larger group than the monocotyledons and the cone-bearing plants combined ([Figures 78-81]).
No matter from what part of the world a totally unfamiliar plant may come, it is always possible to decide into which one of these groups it belongs. That in itself tells us a good deal about its ancestors and its future, “places” it, in fact, in one of those major groups into which all plants are divided. No other characters that plants possess are so important in determining their true position in the scale of plant life as those we have briefly outlined. But merely to sort plants into these large groups does not tell us all we need to know about them. For all plants not only belong to monocotyledons, or dicotyledons, or gymnosperms, but also to smaller divisions of these groups. Just as white men are divided into Englishmen, Frenchmen, etc., so there is the greatest necessity of dividing our large plant groups into smaller and more precise categories.
Some of the chief subdivisions of these large groups have been decided upon the fact that a considerable number of plants in them have some character in common, not found in the remaining plants of the group. Among the monocotyledons, for instance, there is a large class of plants that have tiny flowers between dry, chaffy scales, bear no true petals or sepals, all wind pollinated and are all commonly, though incorrectly, called grasses. These include, strictly speaking, two groups; one, the true grasses in which the stem is mostly hollow and the fruit a grain, while the other, with solid stems and bearing achenes for fruits, are the sedges. The grasses form one family and the sedges another, but while they differ in the characters just mentioned they agree in having flowers of the same general type. Families of plants are thus groups of genera, placed together in the scheme of classification, because they are more like one another than like any other such group. Among the grasses, for instance, are corn, wheat, rice, bamboo, orchard grass, Kentucky blue grass, sugar cane, and hundreds of others, all belonging to different genera, but all those genera grouped into a single family because of their generally similar flowers. Just as the Kentucky blue grass has a generic name (Poa) and a specific one (pratensis), the families of plants must also bear names, usually derived from the generic name of one of the chief genera in it. Because Poa is a large and important genus of the grasses, the family is named after it, with the addition of ceæ. Poaceæ is thus the family name of all the grasses. Among the sedges one of the commonest genera is Cyperus, including many species of the galingale or earth almond. From this genus the sedge family has been named Cyperaceæ ([Figure 87]). So the rose family is the Rosaceæ, the violet family is Violaceæ, and so on through all the three hundred or more families which contain all the flowering plants so far discovered. Going back for a moment to the Poaceæ and Cyperaceæ, the fact that these two large families are different from each other, but have some characters in common, fixes them as both belonging to one order. Orders are thus groups of one or more plant families, all differing one from another, but obviously related and having some characters in common. The order containing the grasses and sedges is named for one of the families in it with the ending ales. Thus Poales include Poaceæ and Cyperaceæ. Rosales include Rosaceæ and several families.
In other words, individual plants are grouped in species, species into genera, genera into families, and families into orders. These orders are themselves grouped into still larger divisions; there are, for instance, twelve orders comprising all the monocotyledons, and about twenty orders comprising all the dicotyledons. Once we have decided that any plant is a monocotyledon or a dicotyledon, our next step should be as to which order it belongs, then its family, its genus, and finally its species. Needless to say, such studies are necessarily of a technical nature, and while the details of them lie outside the scope of this book, the general plan or scheme of flowering plant classification is as we have outlined it above.
This scheme of plant classification has been developed not only for our convenience in sorting plants into definite categories, but more important still to show, if possible, the relationships, and particularly the development from the simplest to the most complex types of plant life. Thus the monocotyledons begin with the cat-tails, which have mere bristles for calyx and corolla, and lead by infinite gradations to the showy and highly complex orchids, which are considered the climax of the monocotyledonous families. While no general account of the plant families can be attempted here, some of the more interesting in both the monocotyledonous and dicotyledonous groups will be briefly discussed.