THE ORGANIC WORLD
(From The Elements of Science.)
By ST. GEORGE MIVART F.R.S.
The number of all the various kinds of living creatures is so enormous that it would be impossible to study them profitably, were they not classified in an orderly manner. Therefore the whole mass has been divided, in the first place, into two supreme groups, fancifully termed kingdoms—the "animal kingdom" and the "vegetal kingdom." Each of these is subdivided into an orderly series of subordinate groups, successively contained one within the other, and named sub-kingdoms, classes, orders, families, genera and species. The lowest group but one is the "genus," which contains one or more different kinds termed "species," as e.g., the species "wood anemone" and the species "blue titmouse." The lowest group of all—a species—may be said to consist of individuals which differ from each other only by trifling characters, such as characters due to difference of sex, while their peculiar organization is faithfully reproduced by generation as a whole, though small individual differences exist in all cases.
The vegetal, or vegetable, kingdom, consists of the great mass of flowering plants, many of which, however, have such inconspicuous flowers that they are mistakenly regarded as flowerless, as is often the case with the grasses, the pines, and the yews. Another mass, or sub-kingdom, of plants consists of the really flowerless plants, such as the ferns, horsetails (Fig. 1), lycopods, and mosses. Sea and fresh-water weeds (algæ), and mushrooms, or "moulds," of all kinds (fungi), amongst which are the now famous "bacteria," constitute a third and lowest set of plants.
The animal kingdom consists, first, of a sub-kingdom of animals which possess a spinal column, or backbone, and which are known as vertebrate animals. Such are all beasts, birds, reptiles, and fishes. There are also a variety of remotely allied marine organisms known as tunicates, sea-squirts, or ascidians (Fig. 2). There is, further, an immense group of arthropods, consisting of all insects, crab-like creatures, hundred-legs and their allies, with spiders, scorpions, tics and mites. We have also the sub-kingdom of shell-fish or molluscs, including cuttle-fishes, snails, whelks, limpets, the oyster, and a multitude of allied forms. A multitudinous sub-kingdom of worms also exists, as well as another of star-fishes and their congeners. There is yet another of zoophytes, or polyps, and another of sponges, and, finally, we have a sub-kingdom of minute creatures, or animalculæ, of very varied forms, which may make up the sub-kingdom of Protozoa, consisting of animals which are mostly unicellular.
Multitudinous and varied as are the creatures which compose this immense organic world, they nevertheless exhibit a very remarkable uniformity of composition in their essential structure. Every living creature from a man to a mushroom, or even to the smallest animalcule or unicellular plant is always partly fluid, but never entirely so. Every living creature also consists in part (and that part is the most active living part) of a soft, viscid, transparent, colorless substance, termed protoplasm, which can be resolved into the four elements, oxygen, hydrogen, nitrogen and carbon. Besides these four elements, living organisms commonly contain sulphur, phosphorus, chlorine, potassium, sodium, calcium, magnesium and iron.
In the fact that living creatures always consist of the four elements, oxygen, hydrogen, nitrogen and carbon, we have a fundamental character whereby the organic and inorganic (or non-living) worlds are to be distinguished, for as we have seen, inorganic bodies, instead of being thus uniformly constituted, may consist of the most diverse elements and sometimes of but two or even of only one.
Again, many minerals, such as crystals, are bounded by plain surfaces, and, with very few exceptions (spathic and hematite iron and dolomite are such exceptions) none are bounded by curved lines and surfaces, while living organisms are bounded by such lines and surfaces.
Yet, again, if a crystal be cut through, its internal structure will be seen to be similar throughout. But if the body of any living creature be divided, it will, at the very least, be seen to consist of a variety of minute distinct particles, called "granules," variously distributed throughout its interior.
All organisms consist either—as do the simplest, mostly microscopic, plants and animals—of a single minute mass of protoplasm, or of a few, or of many, or of an enormous aggregation of such before-mentioned particles, each of which is one of those bodies named a "cell" (Fig. 3). Cells may, or may not, be enclosed in an investing coat or "cell-wall." Every cell generally contains within it a denser, normally spheroidal, body known as the nucleus.
Now protoplasm is a very unstable substance—as we have seen many substances are whereof nitrogen is a component part—and it possesses active properties which are not present in the non-living, or inorganic world. In the latter, differences of temperature will produce motion in the shape of "currents," as we have seen with respect to masses of air and water. But in a portion of protoplasm, an internal circulation of currents in definite lines will establish itself from other causes.
Inorganic bodies, as we have seen, will expand with heat, as they may also do from imbibing moisture; but living protoplasm has an apparently spontaneous power of contraction and expansion under certain external conditions which do not occasion such movements in inorganic matter.
n, nucleus; n', nucleolus embedded in the network of chromatin threads; k, network of the cell external to the nucleus; a, attraction-sphere or archoplasm containing minute bodies called centrosomes; cl, membrane enclosing the cell externally, nl, membrane surrounding the nucleus; c, centrosomes.
Under favoring conditions, protoplasm has a power of performing chemical changes, which result in producing heat far more gently and continuously than it is produced by the combustion of inorganic bodies. Thus it is that the heat is produced which makes its presence evident to us in what we call "warm-blooded animals," the most warm-blooded of all being birds.
Protoplasm has also the wonderful power of transforming certain adjacent substances into material like itself—into its own substance—and so, in a sense, creating a new material. Thus it is that organisms have the power to nourish themselves and grow. An animal would vainly swallow the most nourishing food if the ultimate, protoplasmic particles of its body had not this power of "transforming" suitable substances brought near them in ways to be hereinafter noticed.
Without that, no organism could ever "grow." The growth of organisms is utterly different from the increase in size of inorganic bodies. Crystals, as we have seen, grow merely by external increment; but organisms grow by an increment which takes place in the very innermost substance of the tissues which compose their bodies, and the innermost substance of the cells which compose such tissues; this peculiar form of growth is termed intussusception.
Protoplasm, after thus augmenting its mass, has a further power of spontaneous division, whereby the mass of the entire organism whereof such protoplasm forms a part, is augmented and so growth is brought about.
The small particles of protoplasm which constitute "cells" are far indeed from being structureless. Besides the nucleus already mentioned there is a delicate network of threads of a substance called chromatin within it, and another network permeating the fluid of the cell substance, which invest the nucleus often with further complications. These networks generally perform (or undergo) a most complex series of changes every time a cell spontaneously divides. In certain cases, however, it appears that the nucleus divides into two in a more simple fashion, the rest of the cell contents subsequently dividing—each half enclosing one part of the previously divided nucleus. It is by a continued process of cell division that the complex structures of the most complex organisms is brought about.
The division of a cell, or particle of protoplasm, is indeed a necessary consequence of its complete nutrition.
For new material can only be absorbed by its surface. But as the cell grows, the proportion borne by its surface to its mass, continually decreases; therefore this surface must soon be too small to take in nourishment enough, and the particle, or cell, must therefore either die or divide. By dividing, its parts can continue the nutritive process till their surface, in turn, becomes insufficient, when they must divide again, and so on. Thus the term "feeding" has two senses. "To feed a horse," ordinarily means to give it a certain quantity of hay, oats or what not; and such indeed is one kind of feeding. But obviously, if the nourishment so taken could not get from the stomach and intestines into the ultimate particles and cells of the horse's body, the horse could not be nourished, and still less could it grow. It is this latter process, called assimilation, which is the real and essential process of feeding, to which the process ordinarily so called is but introductory.
Protoplasm has also the power of forming and ejecting from its own substance, other substances which it has made, but which are of a different nature to its own. This function, as before said, is termed secretion; and we know the liver secretes bile, and that the cow's udder secretes milk.
Here again we have an external and an internal process. The milk is drawn forth from a receptacle, the udder, into which it finds its way, and so, in a superficial sense, it may be called an organ of secretion. Nevertheless the true internal secretion takes place in the innermost substance of the cells or particles of protoplasm, of the milk-land, which particles really form that liquid.
FIG. 4. AMOEBA SHOWN IN TWO OF THE MANY IRREGULAR SHAPES IT ASSUMES. (After Howes.)
The clear space within it is a contractile vesicle. The dark body is the nucleus. In the right-hand figure there is shown a particle of food, passing through the external surface.
But every living creature consists at first entirely of a particle of protoplasm. Therefore every other kind of substance which may be found in every kind of plant or animal, must have been formed through it, and be, in fact, a secretion from protoplasm. Such is the rosy cheek of an apple, or of a maiden, the luscious juice of the peach, the produce of the castor-oil plant, the baleen that lines the whale's enormous jaws, as well as that softest product, the fur of the chinchilla. Indeed, every particle of protoplasm requires, in order that it may live, a continuous process of exchange. It needs to be continuously first built up by food, and then broken down by discharging what is no longer needful for its healthy existence. Thus the life of every organism is a life of almost incessant change, not only in its being as a whole, but in that of all its protoplasmic particles also.
Prominent among such processes is that of an interchange of gases between the living being and its environment. This process consists in an absorption of oxygen and a giving-out of carbonic acid, which exchange is termed respiration.
Lastly, protoplasm has a power of motion when appropriately acted on. It will then contract or expand its shape by alternate protrusions and retractions of parts of its substance. These movements are termed amoebiform, because they quite resemble the movements of a small animalcule which is named amoeba. (See Fig. 4.)
Such is the ultimate structure, and such are the fundamental activities or functions of living organisms, as far as they can here be described, from the lowest animalcule and unicellular plant, up to the most complex organisms and the body of man himself.
