Certain species of Pecopteris fronds from Carboniferous strata are characterised by circular sori or synangia consisting of a small number (3–8) of exannulate sporangia attached to a central receptacle and free only at their apices. Strasburger[997] suggested a Marattiaceous affinity for Asterotheca and Stur[998] describes the species Asterotheca Sternbergii Goepp. ([fig. 291], C, D) as an example of a Marattiaceous fern. The latter author retains Corda’s genus Hawlea[999] for the fertile fronds of the common Coal-Measures species Pecopteris Miltoni, while on the other hand Kidston[1000] includes this type in Asterotheca.

Pecopteris (Asterotheca) Miltoni (Artis).
1825.Filicites Miltoni, Artis, Antedil. Phyt. Pl. XIV.
1828.Pecopteris Miltoni, Brongniart, Prodrome, p. 58.
1828.Pecopteris abbreviata, Brongniart, Hist. vég. foss. p. 337, Pl. CXV. figs. 1–4; Lindley and Hutton, Foss. Flor. Vol. III. Pl. 184.
1845.Hawlea pulcherrima, Corda, Flor. Vorwelt, p. 90, Pl. LVII. figs. 7, 8.
1877–1888.Hawlea Miltoni, Stur, Culm Flora, p. 293; Farne Carbon. Flora, p. 108, Pls. LIX. LX.
1888.Pecopteris (Asterotheca) abbreviata, Zeiller, Flor. Valenc. p. 186, Pl. XXIV. figs. 1–4.

Fig. 290.

  1. Alethopteris lonchitica. × 2½. | For description
  2. Lonchopteris rugosa. × 2.| see; Chap. XXVII.
  3. Sphenopteris Hoeninghausi. × 4.|
  4. Parapecopteris neuropteroides.
  5. Pecopteris (Dactylotheca) plumosa [= P. (Dactylotheca) dentata Zeiller (88)]. × 4.

(A–C, E, after Zeiller; D, after Grand’Eury.)

The fronds of this species reached a length of more than 3 metres and a breadth of 2 metres. They are characterised by the presence of aphlebiae[1001] appressed to the rachis and by circular sori composed of a small number (3–6) of sporangia. In habit and in the form of the pinnules this type is similar to Dactylotheca plumosa.

Fig. 291.