The problem of alimentation may be looked at in a thousand ways. It is culinary, no doubt, and gastronomic; but it is also economical and social, agricultural, fiscal, hygienic, medical, and even moral. But first and foremost, it is physiological. It comprises and assumes the knowledge of the general composition of foods, of their transformations in the digestive apparatus, and their comparative utility in the maintenance and the sound functional activity of the organism. To this first group of subjects for our discussion are attached others relating to the effects of inanition, of insufficient alimentation, and of over-feeding. And in order to throw light on all these aspects of the problem of alimentation, we have to lay bare the most intimate and delicate reactions by which the organism is maintained and recruited, and, in the words of a celebrated physiologist, “to penetrate into the kitchen of vital phenomena.” And here neither Apicius, nor Brillat-Savarin, nor Berchoux, nor the moralists, nor the economists are of any use to us as guides. We must appeal to the scientists, who, following the example of Lavoisier, Berzelius, Regnault, and Liebig, have applied to the study of living beings the resources of general science, and have thus founded chemical biology.
This branch of science developed considerably in the second half of the nineteenth century. It has now its methods, its technique, its chairs at the universities, its laboratories, and its literature. It has particularly applied itself to the study of the “material changes” or the metabolism of living beings, and with that object in view it has done two things. In the first place, it has determined the composition of the constituent materials of the organism; then analyzing qualitatively and quantitatively all that penetrates into that organism in a given time—that is to say, all the alimentary or respiratory ingesta, and all that issues from the organism, i.e., all the excreta, all the egesta,—it has drawn up nutritive balance sheets, corresponding to the various conditions of life, whether naturally or artificially created. And thus we can determine the alimentary régimes which give too much, and which give too little, and which finally restore equilibrium.
We do not propose to give a detailed account of this scientific movement. This may be done in monographs. All we wish to indicate here is the most general result of these laborious researches—that is to say, the laws and the doctrines which are derived from them, and the theories to which they have given birth. It is by this alone that they are brought into relation with general science, and may therefore interest the reader. The facts of detail are never lacking to the historian; it is more profitable to show the movement of ideas. The theories of alimentation bring into conflict very different conceptions of the vital functional activity. And here we find a confused medley of opinions on which it is not without interest to endeavour to throw some light.
§ 1. Food, a Source of Energy and Matter.
Definitions of Food.—Before the introduction into physiology of the notion of energy, no one had succeeded in giving an exact idea and a precise definition of food and alimentation. Every physiologist and medical man who attempted it had failed, and this for various reasons.
The general cause of this failure was that most definitions, popular or technical, interposed the condition that the food must be introduced into the digestive apparatus. “It is,” said they, “a substance which when introduced into the digestive tube undergoes, etc., etc.” But plants draw food from the soil, and they possess no digestive apparatus; many animals have no intestinal tube; and in the case of certain rotifera, the females possess a digestive apparatus, while the males have none. Nevertheless all animals feed.
On the other hand, there are other substances than those which use the digestive tract for the purpose of entering the organism, and which are eminently useful or necessary to the maintenance of life. In particular we may mention oxygen.
The distinctive feature of food is its utility—when conveniently introduced or employed—to the living being. Claude Bernard’s definition is this:—A substance taken in the external medium “necessary for the maintenance of the phenomena of the healthy organism and for the reparation of the losses it constantly suffers.” “A substance which supplies an element necessary for the constitution of the organism, or which diminishes its disintegration” (stored-up food); this is the definition of C. Voit, the German physiologist. M. Duclaux says, in his turn, but in far too general terms, that it is a substance which contributes to assure the sound functional activity of any of the organs of the living being. None of these ways of describing food gives a complete idea.
Food, the Source of Energy and Matter.—The intervention of the notion of energy enables us more completely to understand the true nature of food. We must, in fact, have recourse to the energetic conception if we desire to take into account all that the organism requires from food. It not only requires matter, but also, and most important of all, energy.
Investigators so far concentrated their thoughts exclusively on the necessity of a supply of matter—that is to say, they only looked upon one side of the problem. The living body presents, at each of its points, an uninterrupted series of disintegrations and reconstitutions, the materials being supplied from without by alimentation, and rejected by excretion. Cuvier gave to this unceasing circulation of ambient matter throughout the vital world the name of vital vortex, and he rightly saw in it the characteristic of nutrition, and the distinctive feature of life.