CHAPTER II

REFLEX ACTION AND THE FUNCTIONS OF THE SPINAL CORD

I

Until 1820 physiologists believed that all nerves had the same functions; that is, that all were sensory.

We can scarcely picture the confusion in the mind of anyone studying the nerves of the face when, besides those destined to the organs of smell, sight, and hearing, he would notice two other large nerves—the Trigeminus and Facialis—passing off separately from the brain and spinal marrow, and which, with a double ramification of filaments, cover all superficial and underlying parts of the face; and again, when he saw the three nerves of various origin which go to the tongue, the four which are distributed in the throat, and finally, in the midst of this net of nerves, thick bundles of fine filaments and ganglia of which the origin was untraceable.

It was an English physiologist, Charles Bell, who solved this problem by showing that the most important nerves of the face, with the exception of the special sensory nerves, are confined to two. If one of these nerves, called the trigeminal, be cut through, every trace of sensibility immediately disappears from the corresponding side of the face; if the other, the facial nerve, be severed, sensibility remains, but the face completely loses the power of movement, there is no longer any contraction of the muscles or change of expression in the face.

I quote Charles Bell’s own words, since these two simple experiments still form the base of the physiology of the nervous system.

'If we cut the division of the fifth nerve which goes to the lips of an ass, we deprive the lips of sensibility; so when the animal presses the lips to the ground, and against the oats lying there, it does not feel them, and consequently there is no effort made to gather them. If, on the other hand, we cut the seventh nerve where it goes to the lips, the animal feels the oats, but it can make no effort to gather them, the power of muscular motion being cut off by the division of the nerve.’[4]

The same takes place in the hand, the legs, and in all other parts of the body, which, according as the one or the other set of nerves is injured, feel but cannot move, or move and do not feel.