Growing on old prostrate trunks. Æthalium two or three to several centimeters in extent, the individual sporangia 3–5 mm. in height. Plasmodium white, the immature sporangia dull-gray tinged with sienna color. The columella, with its radiating bits of membrane, is the same substance as the wall; it may be a reëntrant edge of the prismatic sporangium, caused by excessive crowding together; at least, this may be regarded as its origin; there may have arisen some further adaptation. The species is Siphoptychium Casparyi, Rost. I am indebted to Dr. George A. Rex for the specimens I have examined.
3. Tubulina cæspitosa, Peck. Sporangia short-cylindric, closely crowded, distinct or connate, argillaceous olive to olive-brown in color, seated on a well-developed hypothallus; the wall a thin membrane, with a dense layer of minute dark-colored round granules on the inner surface. Spores argillaceous olive in the mass, globose, minutely warted, 6–8 mic. in diameter.
Growing on old wood. Æthalium in irregular patches sometimes several centimeters in extent, the single sporangia about 1 mm. in height. Plasmodium dark olivaceous, the sporangia blackish if dried when immature, taking a paler shade of olivaceous, according to development and maturity. This is Perichæna cæspitosa, Peck, in the 31st N. Y. Report.
III. LYCOGALA. Mich. Æthalium with a firm membranaceous wall; from the inner surface of the wall proceed numerous slender tubules, which are intermingled with the spores. The material of the wall appears under three different forms: the inner layer is a thin membrane, uniform in structure, of a yellow-brown color, and semi-pellucid; the outer layer consists of large flat roundish or irregular vesicles, brown in color, filled with minute granules, and arranged in one or more strata; from these vesicles originate the tubules, which traverse the wall for a certain distance, and then enter the interior among the spores; the tubules are more or less compressed, simple or branched, and the surface is ornamented with warts and ridges, which sometimes form irregular rings and reticulations.
If the sporophores in this genus be regarded as simple sporangia, which is the view that Rostafinski takes of one of the species, the tubules are simply the peculiar threads of a capillitium. If, however, the æthalium is a compound plasmodiocarp, the tubules stand for the original plasmodial strands and, consequently, represent the component sporangia.
1. Lycogala conicum. Pers. Æthalia small, ovoid-conic, gregarious, sometimes close together with the bases confluent, the surface pale umber or olivaceous marked with short brown lines, regularly dehiscent at the apex. The wall thin; the outer layer not continuous, the irregular brown vesicles disposed in angular patches and elongated bands, which have a somewhat reticulate arrangement. The tubules appear as a thin stratum upon the inner membrane; they do not branch, and they send long slender simple extremities inward among the spores. Spores in mass pale ochraceous, globose, minutely warted, 5–6 mic. in diameter. [See Plate III, Fig. 5.]
Growing on old wood. Æthalium 2–5 mm. in height, the tubules 3–8 mic. in thickness. This is Dermodium conicum of Rostafinski's monograph, but the structure is essentially the same as in the other species. Massee evidently did not have specimens of this species. I have never seen any branching of the tubules either in the wall or in the free extremities of the interior.
2. Lycogala exiguum, Morg. n. sp. Æthalia small, globose, gregarious, the surface dark brown or blackish, minutely scaly, irregularly dehiscent. The wall thin; the vesicles with a dark polygonal outline, disposed in thin irregular reticulate patches, which are more or less confluent. The tubules appear as an interwoven fibrous stratum upon the inner membrane; they send long slender branched extremities inward among the spores. Spores in mass pale ochraceous, globose, nearly smooth, 5–6 mic. in diameter. [See Plate III, Fig. 6.]
Growing on old wood. Æthalium 2–5 mm. in diameter, the threads 2–10 mic. in thickness, with very slight thickenings of the membrane. The polygonal vesicles give a reticulate appearance to the dark-brown patches which ornament the surface of the wall.
3. Lycogala epidendrum, Buxb. Æthalia subglobose, gregarious, sometimes closely crowded and irregular, the surface umber, brown or olivaceous, minutely warted, at length, irregularly dehiscent at or about the apex. The wall thick, the brown vesicles loosely aggregated and densely agglutinated together, traversed in all directions by the much-branched tubules, which send long-branched extremities inward among the spores; the main branches thick and flat, with wide expansions, especially at the angles, the ultimate branchlets more slender and obtuse at the apex. Spores in the mass from pale to reddish ochre, globose, minutely warted, 5–6 mic. in diameter. [See Plate III, Fig. 7.]