In the course of this introductory sketch we shall find it necessary to mention the names of some of the Orders of Insects that will only be explained or defined in subsequent pages. We may therefore here state that the term "Orthoptera" includes grasshoppers, locusts, earwigs, cockroaches; "Neuroptera" comprises dragon-flies, May-flies, lacewings, stone-flies and caddis-flies; to the "Hymenoptera" belong bees, wasps, ants, sawflies, and a host of little creatures scarcely noticed by the ordinary observer: "Coleoptera" are beetles; "Lepidoptera," butterflies and moths; "Diptera," house-flies, blue-bottles, daddy-longlegs, and such; "Hemiptera" or "Rhynchota" are bugs, greenfly, etc.

Class Insecta: or Insecta Hexapoda.

Definition.—Insects are small animals, having the body divided into three regions placed in longitudinal succession—head, thorax, and abdomen: they take in air by means of tracheae, a system of tubes distributed throughout the body, and opening externally by means of orifices placed at the sides of the body. They have six legs, and a pair of antennae; these latter are placed on the head, while the legs are attached to the thorax, or second of the three great body divisions; the abdomen has no true legs, but not infrequently has terminal appendages and, on the under surface, protuberances which serve as feet. Very frequently there are two pairs of wings, sometimes only one pair, in other cases none: the wings are always placed on the thorax. Insects are transversely segmented—that is to say, the body has the form of a succession of rings; but this condition is in many cases obscure; the number of these rings rarely, if ever, exceeds thirteen in addition to the head and to a terminal piece that sometimes exists. Insects usually change much in appearance in the course of their growth, the annulose or ringed condition being most evident in the early part of the individual's life. The legs are usually elongate and apparently jointed, but in the immature condition may be altogether absent, or very short; in the latter case the jointing is obscure. The number of jointed legs is always six.

External Structure.

The series of rings of which the external crust or skeleton of Insects is composed exhibits great modifications, not only in the various kinds of Insects but even in the different parts of the same individual, and at successive periods of its development; so that in the majority of mature Insects the separate rings are readily distinguished only in the hind body or abdomen. The total number of the visible rings, segments, somites, or arthromeres, as they are variously called by different writers, is frequently thirteen in addition to the head. This latter part is considered to be itself composed of the elements of several rings, but morphologists are not yet agreed as to their number, some thinking this is three while others place it as high as seven; three or four being, perhaps, the figures at present most in favour, though Viallanes, who has recently discussed[[17]] the subject, considers six, the number suggested by Huxley, as the most probable. Cholodkovsky is of a similar opinion. However this may be, the three rings behind the head constitute the thorax, which is always largely developed, though, like the head, its segmentation is usually very much obscured by unequal development of different parts, or by consolidation of some of them, or by both of these conditions. The third great division of the body, the abdomen, is also usually much modified by one or more of the terminal segments being changed in form, or even entirely withdrawn into the interior of the body. The existence of ten segments in the hind body can, however, be very frequently actually demonstrated, so that it is correct to speak of ten as the normal number.

Fig. 47—Diagram of exterior of insect: the two vertical dotted lines indicate the divisions between H, head; T, thorax; and A, abdomen: a, antenna; b, labrum; c, mandible; d, maxillary palpus; e, labial palpus; f, facetted eye; g, pronotum; h, mesonotum; i, metanotum; k, wings; l1 to l10, abdominal segments; m, the internal membranous portions uniting the apparently separated segments; n, cerci; o, stigma; p, abdominal pleuron bearing small stigmata; q1, q2, q3, pro-, meso-, meta-sterna; r1, mesothoracic episternum; s1, epimeron, these two forming the mesopleuron; r2, s2, metathoracic episternum and epimeron; t, coxa; v, trochanter; w, femur; x, tibia; y, tarsus; z, gula.

It is no reproach to morphologists that they have not yet agreed as to the number of segments that may be taken as typical for an Insect, for all the branches of evidence bearing on the point are still imperfect. It may be well, therefore, to state the most extreme views that appear to be at all admissible. Hagen[[18]] has recently stated the opinion that each thoracic segment consists really of three segments—an anterior or wing-bearer, a middle or leg-bearer, and a posterior or stigma-bearer. There seems to be no reason for treating the stigma as being at all of the nature of an appendage, and the theory of a triple origin for these segments may be dismissed. There are, however, several facts that indicate a duplicity in these somites, among which we may specially mention the remarkable constancy of two pleural pieces on each side of each thoracic segment. The hypothesis of these rings being each the representative of two segments cannot therefore be at present considered entirely untenable, and in that case the maximum and minimum numbers that can be suggested appear to be twenty-four and eleven, distributed as follows:—

Maximum.Minimum.
Head73
Thorax63
Abdomen115
Total2411

Although it is not probable that ultimately so great a difference as these figures indicate will be found to prevail, it is certainly at present premature to say that all Insects are made up of the same number of primary segments.