But if the unassisted eye sees life distributed throughout the atmosphere, when armed with the microscope we discover far other marvels. Rotiferæ, Brachionæ, and a multitude of microscopic animalculæ, are carried up by the winds from the surface of evaporating waters. These minute creatures, motionless and apparently dead, are borne to and fro in the air until the falling dews bring them back to the surface of the earth, dissolve the film or envelope which encloses their transparent rotating bodies,[3] and, probably by means of the oxygen which all waters contain, breathe new irritability into their dormant organs.
According to Ehrenberg’s brilliant discovery, the yellow sand or dust which falls like rain on the Atlantic near the Cape de Verde Islands, and is occasionally carried even to Italy and Middle Europe, consists of a multitude of siliceous-shelled microscopic animals. Perhaps many of them float for years in the upper strata of the atmosphere, until they are brought down by vertical currents or in accompaniment with the superior current of the trade-winds, still susceptible of revivification, and multiplying their species by spontaneous division in conformity with the particular laws of their organisation.
But, besides creatures fully formed, the atmosphere contains innumerable germs of future life, such as the eggs of insects and the seeds of plants, the latter provided with light hairy or feathery appendages, by means of which they are wafted through the air during long autumnal wanderings. Even the fertilizing dust or pollen from the anthers of the male flowers, in species in which the sexes are separated, is carried over land and sea, by winds and by the agency of winged insects,[4] to the solitary female plant on other shores. Thus wherever the glance of the inquirer into Nature penetrates, he sees the continual dissemination of life, either fully formed or in the germ.
If the aereal ocean in which we are submerged, and above the surface of which we cannot rise, be indispensable to the existence of organised beings, they also require a more substantial aliment, which they can find only at the bottom of this gaseous ocean. This bottom is of two kinds; the smaller portion consisting of dry land in immediate contact with the external atmosphere, and the larger portion consisting of water, which may perhaps have been formed thousands of years ago by electric agencies from gaseous substances, and which is now incessantly undergoing decomposition in the laboratories of Nature, in the clouds and in the pulsating vessels of animals and plants. Organic forms also descend deep below the surface of the earth, wherever rain or surface water can percolate either by natural cavities or by mines or other excavations made by man: the subterranean cryptogamic Flora was an object of my scientific research in the early part of my life. Thermal springs of very high temperature nourish small Hydropores, Confervæ, and Oscillatoria. At Bear Lake, near the Arctic Circle, Richardson saw the ground, which continues frozen throughout the summer at a depth of twenty inches, covered with flowering plants.
We do not yet know where life is most abundant,—whether on continents or in the unfathomed depths of the ocean. Through the excellent work of Ehrenberg, “Über das Verhalten des kleinsten Lebens,” we have seen the sphere of organic life extend, and its horizon widen before our eyes, both in the tropical parts of the ocean and in the fixed or floating masses of ice of the Antarctic seas. Siliceous-shelled Polygastrica, and even Coscinodiscæ, with their green ovaries, have been found alive enveloped in masses of ice only twelve degrees from the Pole; the small black Glacier flea (Desoria glacialis) and Podurellæ inhabit the narrow tubular holes examined by Agassiz in the Swiss glaciers. Ehrenberg has shown that on several microscopic Infusoria (Synedra, Cocconeis) others live as parasites, and that in the Gallionellæ such is their prodigious power of development, or capability of division, that in the space of four days an animalcule invisible to the naked eye can form two cubic feet of the Bilin polishing slate. In the sea, gelatinous worms, living or dead, shine like stars,[5] and by their phosphoric light change the surface of the wide ocean into a sea of fire. Ineffaceable is the impression made on my mind by the calm nights of the torrid zone, on the waters of the Pacific. I still see the dark azure of the firmament, the constellation of the Ship near the zenith, and that of the Cross declining towards the horizon, shedding through the perfumed air their soft and planetary lustre; while bright furrows of flashing light marked the track of the dolphins through the midst of the foaming waves.
Not only the ocean, but also the waters of our marshes, hide from us an innumerable multitude of strange forms. The naked eye can with difficulty distinguish the Cyclidias, the Euglenes, and the host of Naids divisible by branches like the Lemna or Duckweed, of which they seek the shade. Other creatures inhabit receptacles where the light cannot penetrate, and an atmosphere variously composed, but differing from that which we breathe: such are the spotted Ascaris, which lives beneath the skin of the earthworm; the Leucophra, of a bright silvery colour, in the interior of the shore Naid; and a Pentastoma, which inhabits the large pulmonary cells of the rattlesnake of the tropics.[6] There are animalculæ in the blood of frogs and of salmon, and even, according to Nordmann, in the fluids of the eyes of fishes and in the gills of the Bleak. Thus the most hidden recesses of creation teem with life. We propose in these pages to direct our attention to the vegetable world, on the existence of which that of animals is dependent. Plants are incessantly engaged in disposing into order towards subsequent organization the raw materials of which the earth is composed: it is their office, by their vital forces or powers, to prepare those substances which, after undergoing a thousand modifications, are gradually converted to nobler purposes in the formation of nervous tissues. In directing our consideration towards the various families of plants, we shall at the same time glance at the multitude of animated beings to which they afford nutriment and protection.
The carpet of flowers and of verdure spread over the naked crust of our planet is unequally woven; it is thicker where the sun rises high in the ever cloudless heavens, and thinner towards the poles, in the less happy climes where returning frosts often destroy the opening buds of spring, or the ripening fruits of autumn. Everywhere, however, man finds some plants to minister to his support and enjoyment. If new lands are formed, the organic forces are ever ready to cover the naked rock with life. Sometimes, as at an early period among the Greek Islands, volcanic forces suddenly elevate above the surface of the boiling waves a rock covered with Scoriæ: sometimes, by a long-continued and more tranquil series of phenomena, the collective labours of united Lithophytes[7] raise their cellular dwellings on the crests of submarine mountains, until, after thousands of years, the structure reaches the level of the ocean, when the creatures which have formed it die, leaving a low flat coral island. How are the seeds of plants brought so immediately to these new shores? by wandering birds, or by the winds and waves of the ocean? The distance from other coasts makes it difficult to determine this question; but, no sooner is the rock of the newly raised islands in direct contact with the atmosphere, than there is formed on its surface, in our northern countries, a soft silky net-work, appearing to the naked eye as coloured spots and patches. Some of these patches are bordered by single or double raised lines running round their margins; other patches are crossed by similar lines traversing them in various directions. Gradually the light colour of the patches becomes darker, the bright yellow which was visible at a distance changes to brown, and the bluish gray of the Leprarias becomes a dusty black. The edges of neighbouring patches approach and run into each other; and on the dark ground thus formed there appear other lichens, of a circular shape and dazzling whiteness. Thus an organic film or covering establishes itself by successive layers; and as mankind, in forming settled communities, pass through different stages of civilisation, so is the gradual propagation and extension of plants connected with determinate physical laws. Lichens form the first covering of the naked rock, where afterwards lofty forest trees rear their airy summits. The successive growth of mosses, grasses, herbaceous plants, and shrubs or bushes, occupies the intervening period of long but undetermined duration. The part which lichens and mosses perform in the northern countries is effected within the tropics by Portulacas, Gomphrenas, and other low and succulent shore plants. The history of the vegetable covering of our planet, and its gradual propagation over the desert crust of the earth, has its epochs, as well as that of the migrations of the animal world.
Yet although organic life is everywhere diffused, and the organic powers are incessantly at work in reconnecting with each other the elements set free by death or dissolution, the abundance and variety of organised beings, and the rapidity with which they are renewed, differ in different climates. In the cold zones, the activity of organic life undergoes a temporary suspension during a portion of the year by frost; fluidity is an essential condition of life or vital action, and animals and plants, with the exception of mosses and other cryptogamia, are in those regions buried for several months of each year in winter sleep. Over a large part of the earth, therefore, there could only be developed organic forms capable of supporting either a considerable diminution of heat, or, being without leaves, a long interruption of the vital functions. Thus we see variety and grace of form, mixture of colours, and generally the perpetually youthful energy and vigour of organic life, increase as we approach the tropics. This increase can be denied only by those who have never quitted Europe, or who have neglected the study of physical geography. When, leaving our oak forests, we traverse the Alps or the Pyrenees, and enter Italy or Spain, or when we direct our attention to some of the African shores of the Mediterranean, we might easily be led to draw the erroneous inference that hot countries are marked by the absence of trees. But those who do so, forget that the South of Europe wore a different aspect on the first arrival of Pelasgian or Carthaginian colonies; they forget that an ancient civilisation causes the forests to recede more and more, and that the wants and restless activity of large communities of men gradually despoil the face of the earth of the refreshing shades which still rejoice the eye in Northern and Middle Europe, and which, even more than any historic documents, prove the recent date and youthful age of our civilization. The great catastrophe which occasioned the formation of the Mediterranean, when the swollen waters of what was previously an immense lake burst through the barriers of the Dardanelles and of the Pillars of Hercules, appears to have stripped the adjacent countries of a large portion of their coating of vegetable mould. The traditions of Samothrace,[8] handed down to us by Grecian writers, appear to indicate the recentness of the epoch of the ravages caused by this great change. In all the countries which surround the Mediterranean, and which are characterised by beds of the tertiary and cretaceous periods (nummulitic limestone and neocomian rocks), great part of the surface of the earth consists of naked rock. One especial cause of the picturesque beauty of Italian scenery is the contrast thus afforded between the bare rock, and the islands if I may so call them of luxuriant vegetation scattered over its surface. Wherever the rock is less intersected with fissures, so that it retains water at the surface, and where it is covered with vegetable mould, there, as on the enchanting shores of the Lake of Albano, Italy has her oak forests, with glades as deeply embowered and verdure as fresh as those which we admire in the North of Europe.
The deserts to the south of the Atlas, and the immense plains or steppes of South America, must be regarded as only local phenomena. The latter, the South American steppes, are clothed, in the rainy season at least, with grass, and with low-growing almost herbaceous mimosas. The African deserts are, indeed, at all seasons devoid of vegetation; seas of sand, surrounded by forest shores clothed with perpetual verdure. A few scattered fan-palms alone recall to the wanderer’s recollection that these awful solitudes belong to the domain of the same animated terrestrial creation which is elsewhere so rich and so varied. The fantastic play of the mirage, occasioned by the effects of radiant heat, sometimes causes these palm trees to appear divided from the ground and hovering above its surface, and sometimes shews their inverted image reflected in strata of air undulating like the waves of the sea. On the west of the great Peruvian chain of the Andes, on the coasts of the Pacific, I have passed entire weeks in traversing similar deserts destitute of water.
The origin of extensive arid tracts destitute of plants, in the midst of countries rich in luxuriant vegetation, is a geognostical problem which has hitherto been but little considered, but which has doubtless depended on ancient revolutions of nature, such as inundations or great volcanic changes. When once a region has lost the covering of plants with which it was invested, if the sands are loose and mobile and are destitute of springs, and if the heated atmosphere, forming constantly ascending currents, prevents precipitation taking place from clouds[9], thousands of years may elapse ere organic life can pass from the verdant shores to the interior of the sandy sea, and repossess itself of the domain from which it had been banished.