Pen-y-bryn, St. Peter's Road, Croydon. January 1, 1881.
My dear Darwin,—I have been intending to write to you for some weeks to call your attention to what seems to me a striking confirmation (or at all events a support) of my views of the land migration of plants from mountain to mountain. In Nature of Dec. 9th, p. 126, Mr. Baker, of Kew, describes a number of the alpine plants of Madagascar as being identical species with some found on the mountains of Abyssinia, the Cameroons, and other African [pg 312]mountains. Now, if there is one thing more clear than another it is that Madagascar has been separated from Africa since the Miocene (probably the early Miocene) epoch. These plants must therefore have reached the island either since then, in which case they certainly must have passed through the air for long distances, or at the time of the union. But the Miocene and Eocene periods were certainly warm, and these alpine plants could hardly have migrated over tropical forest lands, while it is very improbable that if they had been isolated at so remote a period, exposed to such distinct climatal and organic environments as in Madagascar and Abyssinia, they would have in both places retained their specific characters unchanged. The presumption is, therefore, that they are comparatively recent immigrants, and if so must have passed across the sea from mountain to mountain, for the richness and speciality of the Madagascar forest vegetation render it certain that no recent glacial epoch has seriously affected that island.
Hoping that you are in good health, and wishing you the compliments of the season, I remain yours very faithfully,
ALFRED R. WALLACE.
Down, Beckenham, Kent. January 2, 1881.
My dear Wallace,—The case which you give is a very striking one, and I had overlooked it in Nature.[116] But I remain as great a heretic as ever. Any supposition seems to me more probable than that the seeds of plants should have been blown from the mountains of Abyssinia or other central mountains of Africa to the mountains of Madagascar. It seems to me almost infinitely more probable [pg 313]that Madagascar extended far to the south during the Glacial period, and that the southern hemisphere was, according to Croll, then more temperate; and that the whole of Africa was then peopled with some temperate forms, which crossed chiefly by agency of birds and sea-currents; and some few by the wind from the shores of Africa to Madagascar, subsequently ascending to the mountains.
How lamentable it is that two men should take such widely different views, with the same facts before them; but this seems to be almost regularly our case, and much do I regret it.
I am fairly well, but always feel half dead with fatigue. I heard but an indifferent account of your health some time ago, but trust that you are now somewhat stronger.—Believe me, my dear Wallace, yours very sincerely,