fruit-eating birds, almost equally abundant in tropical Asia and Africa, but less plentiful in America, where they probably suffer from the competition of the larger sized toucans. The genera of each country are distinct, but all are closely allied, the family being a very natural one. The trogons form a family of very gorgeously coloured and remarkable insect-eating birds very abundant in tropical America, less so in Asia, and with a single genus of two species in Africa.
Among reptiles we have two families of snakes—the Dendrophidæ or tree-snakes, and the Dryiophidæ or green whip-snakes—which are also found in the three tropical regions of Asia, Africa, and America, but in these cases even some of the genera are common to Asia and Africa, or to Africa and America. The lizards forming the family Amphisbænidæ are divided between tropical Africa and America, a few species only occurring in the southern portion of the adjacent temperate regions; while even the peculiarly American family of the iguanas is represented by two genera in Madagascar, and one in the Fiji and Friendly Islands. Passing on to the Amphibians the worm-like Cæciliadæ are tropicopolitan, as are also the toads of the family Engystomatidæ. Insects also furnish some analogous cases, three genera of Cicindelidæ, (Pogonostoma, Ctenostoma, and Peridexia) showing a decided connection between this family in South America and Madagascar; while the beautiful family of diurnal moths, Uraniidæ, is confined to the same two countries. A somewhat similar but better known illustration is afforded by the two genera of ostriches, one confined to Africa and Arabia, the other to the plains of temperate South America.
General features of Overlapping and Discontinuous Areas.—These numerous examples of discontinuous genera and families form an important section of the facts of animal dispersal which any true theory must satisfactorily account for. In greater or less prominence they are to be found all over the world, and in every group of animals, and they grade imperceptibly into those cases of conterminous and overlapping areas which we have seen to
prevail in most extensive groups of species, and which are perhaps even more common in those large families which consist of many closely allied genera. A sufficient proof of the overlapping of generic areas is the occurrence of a number of genera of the same family together. Thus in France or Italy about twenty genera of warblers (Sylviadæ) are found, and as each of the thirty-three genera of this family inhabiting temperate Europe and Asia has a different area, a great number must here overlap. So, in most parts of Africa, at least ten or twelve genera of antelopes may be found, and in South America a large proportion of the genera of monkeys of the family Cebidæ occur in many districts; and still more is this the case with the larger bird families, such as the tanagers, the tyrant shrikes, or the tree-creepers, so that there is in all these extensive families no genus whose area does not overlap that of many others. Then among the moderately extensive families we find a few instances of one or two genera isolated from the rest, as the spectacled bear, Tremarctos, found only in Chili, while the remainder of the family extends from Europe and Asia over North America to the Mountains of Mexico, but no further south; the Bovidæ, or hollow-horned ruminants, which have a few isolated genera in the Rocky Mountains and the islands of Sumatra and Celebes; and from these we pass on to the cases of wide separation already given.
Restricted Areas of Families.—As families sometimes consist of single genera and even single species, they often present examples of very restricted range; but what is perhaps more interesting are those cases in which a family contains numerous species and sometimes even several genera, and yet is confined to a narrow area. Such are the golden moles (Chrysochloridæ) consisting of two genera and three species, confined to extratropical South Africa; the hill-tits (Liotrichidæ), a family of numerous genera and species mainly confined to the Himalayas, but with a few straggling species in the Malay countries and the mountains of China; the Pteroptochidæ, large wren-like birds, consisting of eight genera and nineteen species, almost entirely confined to temperate South America and
the Andes; and the birds-of-paradise, consisting of nineteen or twenty genera and about thirty-five species, almost all inhabitants of New Guinea and the immediately surrounding islands, while a few, doubtfully belonging to the family, extend to East Australia. Among reptiles the most striking case of restriction is that of the rough-tailed burrowing snakes (Uropeltidæ), the five genera and eighteen species being strictly confined to Ceylon and the southern parts of the Indian Peninsula.
The Distribution of Orders.—When we pass to the larger groups, termed orders, comprising several families, we find comparatively few cases of restriction and many of worldwide distribution; and the families of which they are composed are strictly comparable to the genera of which families are composed, inasmuch as they present examples of overlapping, or conterminous, or isolated areas, though the latter are comparatively rare. Among mammalia the Insectivora offer the best example of an order, several of whose families inhabit areas more or less isolated from the rest; while the Marsupialia have six families in Australia, and one, the opossums, far off in America.
Perhaps, more important is the limitation of some entire orders to certain well-defined portions of the globe. Thus the Proboscidea, comprising the single family and genus of the elephants, and the Hyracoidea, that of the Hyrax or Syrian coney, are confined to parts of Africa and Asia; the Marsupials to Australia and America; and the Monotremata, the lowest of all mammals—comprising the duck-billed Platypus and the spiny Echidna, to Australia and New Guinea. Among birds the Struthiones or ostrich tribe are almost confined to the three Southern continents, South America, Africa and Australia; and among Amphibia the tailed Batrachia—the newts and salamanders—are similarly restricted to the northern hemisphere.
These various facts will receive their explanation in a future chapter.