Island Life; Or, The Phenomena and Causes of Insular Faunas and Floras

by Alfred Russel Wallace

ANCIENT GLACIAL EPOCHS, AND MILD CLIMATES IN THE ARCTIC REGIONS

Dr. Croll's Views on Ancient Glacial Epochs—Effects of Denudation in Destroying the Evidence of Remote Glacial Epochs—Rise of Sea-level Connected with Glacial Epochs a Cause of Further Denudation—What Evidence of Early Glacial Epochs may be Expected—Evidences of Ice-action During the Tertiary Period—The Weight of the Negative Evidence—Temperate Climates in the Arctic Regions—The Miocene Arctic Flora—Mild Arctic Climates of the Cretaceous Period—Stratigraphical Evidence of Long-continued Mild Arctic Conditions—The Causes of Mild Arctic Climates—Geographical Conditions Favouring Mild Northern Climates in Tertiary Times—The Indian Ocean as a Source of Heat in Tertiary Times—Condition of North America During the Tertiary Period—Effect of High Excentricity on Warm Polar Climates—Evidences as to Climate in the Secondary and Palæozoic Epochs—Warm Arctic Climates in Early Secondary and Palæozoic Times—Conclusions as to the Climates of Secondary and Tertiary Periods—General View of Geological Climates as Dependent on the Physical Features of the Earth's Surface—Estimate of the Comparative Effects of Geographical and Physical Causes in Producing Changes of Climate.

If we adopt the view set forth in the preceding chapter as to the character of the glacial epoch and of the accompanying alternations of climate, it must have been a very important agent in producing changes in the distribution of animal and vegetable life. The intervening mild periods, which almost certainly occurred during its earlier and later phases, may have been sometimes more equable than even our present insular climate, and severe frosts were probably then unknown. During the four or five

thousand years that each specially mild period may have lasted, some portions of the north temperate zone, which had been buried in snow or ice, would become again clothed with vegetation and stocked with animal life, both of which, as the cold again came on, would be driven southward, or perhaps partially exterminated. Forms usually separated would thus be crowded together, and a struggle for existence would follow, which must have led to the modification or the extinction of many species. When the survivors in the struggle had reached a state of equilibrium, a fresh field would be opened to them by the later ameliorations of climate; the more successful of the survivors would spread and multiply; and after this had gone on for thousands of generations, another change of climate, another southward migration, another struggle of northern and southern forms would take place.

But if the last glacial epoch has coincided with, and has been to a considerable extent caused by, a high excentricity of the earth's orbit, we are naturally led to expect that earlier glacial epochs would have occurred whenever the excentricity was unusually large. Dr. Croll has published tables showing the varying amounts of excentricity for three million years back; and from these it appears that there have been many periods of high excentricity, which has often been far greater than at the time of the last glacial epoch.[^[62]] The accompanying diagram has been drawn from these tables, and it will be seen that the highest excentricity occurred 850,000 years ago, at which time the difference between the sun's distance at aphelion and perihelion was thirteen and a half millions of miles, whereas during the last glacial period the maximum difference was ten and a half million miles.

Now, judging by the amount of organic and physical change that occurred during and since the glacial epoch, and that which has occurred since the Miocene period, it is considered probable that this maximum of excentricity coincided with some part of the latter period; and Dr. Croll maintains that a glacial epoch must then have

occurred surpassing in severity that of which we have such convincing proofs, and consisting like it of alternations of cold and warm phases every 10,500 years. The diagram also shows us another long-continued period of high excentricity from 1,750,000 to 1,950,000 years ago, and yet another almost equal to the maximum 2,500,000 years back. These may perhaps have occurred during the Eocene and Cretaceous epochs respectively, or all may have been included within the limits of the Tertiary period. As two of these high excentricities greatly exceed that which caused our glacial epoch, while the third is almost equal to it and of longer duration, they seem to afford us the means of testing rival theories of the causes of glaciation. If, as Dr. Croll argues, high excentricity is the great and dominating agency in bringing on glacial epochs, geographical changes being subordinate, then there must have been glacial epochs of great severity at all these three periods; while if he is also correct in supposing that the alternate phases of precession would inevitably produce glaciation in one hemisphere, and a proportionately mild and equable climate in the opposite hemisphere, then we should have to look for evidence of exceptionally warm and exceptionally cold periods, occurring

alternately and with several repetitions, within a space of time which, geologically speaking, is very short indeed.

Let us then inquire first into the character of the evidence we should expect to find of such changes of climate, if they have occurred; we shall then be in a better position to estimate at its proper value the evidence that actually exists, and, after giving it due weight, to arrive at some conclusion as to the theory that best explains and harmonises it.