The most recent of these milder climates is perhaps indicated by the abundant remains of large mammalia—such as the mammoth, woolly rhinoceros, bison and horse, in the icy alluvial plains of Northern Siberia, and especially in the Liakhov Islands in the same latitude as the North Cape of Asia. These remains occur not in one or two spots only, as if collected by eddies at the mouth of a river, but along the whole borders of the Arctic Ocean; and it is generally admitted that the animals must have lived upon the adjacent plains, and that a considerably milder climate than now prevails could alone have enabled them to do so. How long ago this occurred we do not know, but one of the last intercalated mild periods of the glacial epoch itself seems to offer all the necessary conditions. Again, Sir Edward Belcher discovered on the dreary shores of Wellington Channel in 75½° N. Lat. the trunk and root of a fir tree which had evidently grown where it was found. It appeared to belong to the species Abies alba, or white fir, which now reaches 68° N. Lat. and is the most northerly conifer known. Similar trees, one four feet in circumference and thirty feet long, were found by Lieut. Mecham in Prince Patrick's Island in Lat. 76° 12′ N., and other Arctic explorers have found remains of trees in high latitudes.[[67]]
Similar indications of a recent milder climate are found in Spitzbergen. Professor Nordenskjöld says: "At various places on Spitzbergen, at the bottom of Lomme Bay, at Cape Thordsen, in Blomstrand's strata in Advent Bay, there are found large and well-developed shells of a bivalve, Mytilus edulis, which is not now found living on the coast of Spitzbergen, though on the west coast of Scandinavia it everywhere covers the rocks near the sea-shore. These shells occur most plentifully in the bed of a river which runs through Reindeer Valley at Cape Thordsen. They
are probably washed out of a thin bed of sand at a height of about twenty or thirty feet above the present sea-level, which is intersected by the river. The geological age of this bed cannot be very great, and it has clearly been formed since the present basin of the Ice Sound, or at least the greater part of it, has been hollowed out by glacial action."[[68]]
The Miocene Arctic Flora.—One of the most startling and important of the scientific discoveries of the last forty years has been that of the relics of a luxuriant Miocene flora in various parts of the Arctic regions. It is a discovery that was totally unexpected, and is even now considered by many men of science to be completely unintelligible; but it is so thoroughly established, and it has such a direct and important bearing on the subjects we are discussing in the present volume, that it is necessary to lay a tolerably complete outline of the facts before our readers.
The Miocene flora of temperate Europe was very like that of Eastern Asia, Japan, and the warmer part of Eastern North America of the present day. It is very richly represented in Switzerland by well preserved fossil remains, and after a close comparison with the flora of other countries Professor Heer concludes that the Swiss Lower Miocene flora indicates a climate corresponding to that of Louisiana, North Africa, and South China, while the Upper Miocene climate of the same country would correspond to that of the south of Spain, Southern Japan, and Georgia (U.S. of America). Of this latter flora, found chiefly at Œninghen in the northern extremity of Switzerland, 465 species are known, of which 166 species are trees or shrubs, half of them being evergreens. They comprise sequoias like the Californian giant trees, camphor-trees, cinnamons, sassafras, bignonias, cassias, gleditschias, tulip-trees, and many other American genera, together with maples, ashes, planes, oaks, poplars, and other familiar European trees represented by a variety of extinct species. If we now go to the west coast of Greenland in 70° N. Lat. we find abundant remains of a flora of the same general
type as that of Œninghen but of a more northern character. We have a sequoia identical with one of the species found at Œninghen, a chestnut, salisburia, liquidambar, sassafras, and even a magnolia. We have also seven species of oaks, two planes, two vines, three beeches, four poplars, two willows, a walnut, a plum, and several shrubs supposed to be evergreens; altogether 137 species, mostly well and abundantly preserved!
But even further north, in Spitzbergen, in 78° and 79° N. Lat. and one of the most barren and inhospitable regions on the globe, an almost equally rich fossil flora has been discovered including several of the Greenland species, and others peculiar, but mostly of the same genera. There seem to be no evergreens here except coniferæ, one of which is identical with the swamp-cypress (Taxodium distichum) now found living in the Southern United States! There are also eleven pines, two Libocedrus, two sequoias, with oaks, poplars, birches, planes, limes, a hazel, an ash, and a walnut; also water-lilies, pond-weeds, and an iris—altogether about a hundred species of flowering plants. Even in Grinnell Land, within 8¼ degrees of the pole, a similar flora existed, twenty-five species of fossil plants having been collected by the last Arctic expedition, of which eighteen were identical with the species from other Arctic localities. This flora comprised poplars, birches, hazels, elms, viburnums, and eight species of conifers including the swamp cypress and the Norway spruce (Pinus abies) which last does not now extend beyond 69½° N.
Fossil plants closely resembling those just mentioned have been found at many other Arctic localities, especially in Iceland, on the Mackenzie River in 65° N. Lat. and in Alaska. As an intermediate station we have, in the neighbourhood of Dantzic in Lat. 55° N., a similar flora, with the swamp-cypress, sequoias, oaks, poplars, and some cinnamons, laurels, and figs. A little further south, near Breslau, north of the Carpathians, a rich flora has been found allied to that of Œninghen, but wanting in some of the more tropical forms. Again, in the Isle of Mull in Scotland, in about 56½° N. Lat., a plant-bed has been discovered
containing a hazel, a plane, and a sequoia, apparently identical with a Swiss Miocene species.
We thus find one well-marked type of vegetation spread from Switzerland and Vienna to North Germany, Scotland, Iceland, Greenland, Alaska, and Spitzbergen, some few of the species even ranging over the extremes of latitude between Œninghen and Spitzbergen, but the great majority being distinct, and exhibiting decided indications of a decrease of temperature according to latitude, though much less in amount than now exists. Some writers have thought that the great similarity of the floras of Greenland and Œninghen is a proof that they were not contemporaneous, but successive; and that of Greenland has been supposed to be as old as the Eocene. But the arguments yet adduced do not seem to prove such a difference of age, because there is only that amount of specific and generic diversity between the two which might be produced by distance and difference of temperature, under the exceptionally equable climate of the period. We have even now examples of an equally wide range of well-marked types; as in temperate South America, where many of the genera and some of the species range from the Straits of Magellan to Valparaiso—places differing as much in latitude as Switzerland and West Greenland; and the same may be said of North Australia and Tasmania, where, at a greater latitudinal distance apart, closely allied forms of Eucalyptus, Acacia, Casuarina, Stylidium, Goodenia, and many other genera would certainly form a prominent feature in any fossil flora now being preserved.