Each of these nervous masses contains nerve-cells as well as nerve-fibres, and is capable of generating nerve-force. Each, therefore, acts like a minute brain; and, in fact, the entire ganglionic system of nerves is analogous to the nervous system of certain among the lower animals—the crustacea and mollusks. These possess neither brain nor spinal cord; their nerve-centres, instead of being concentrated in a cranium and vertebral canal, are entirely disseminated through the cavities of the trunk, as are the visceral plexuses in vertebrated animals. In these, however, the addition of a brain and spinal cord to the original rudimentary nervous system, powerfully modifies and controls the action of the latter. The degree of control is variable, according to the relative predominance of the one or the other; and this predominance varies, not only according to different species of vertebrated animals, but also according to different individuals, in that which presents the most conspicuous capacity for individual variation—the human species. Up to a certain point, increased development of the cerebro-spinal system, attended by an increased development of the osseo-muscular framework of the body, is also accompanied by greater elaboration of the ganglionic nerves supplying the viscera, upon whose efficient action the nutrition of this frame depends. But beyond a certain point in the ascending scale, the exactness of this correlation ceases. The muscles and bones are smaller; yet the structure of the cerebro-spinal organs, especially the brain, becomes more elaborate; and hence the control exercised over the functions of the ganglionic system is more complete, although the relative size of the two systems is not much changed.

Such control or predominance is manifested in the following ways: First. The functions of animal life, presided over by the cerebro-spinal system, become proportionately more important than those of vegetative or nutritive life, carried on by the ganglionic. That is to say, the acts of locomotion sustained by the spinal cord and the nervo-muscular apparatus, and the intellectual acts of the thought and will, sustained by the brain—are relatively more prominent than are the acts of digestion, respiration, circulation, etc., dependent on the functions of the ganglionic nerves. Second. These latter functions are themselves effected with more regularity and more force, when the activity of the cerebro-spinal system predominates over that of the ganglionic. Within certain limits, this is so true, that human beings possess over lower animals a superiority, not only of intellect, but of capacity for digesting various articles of food; and of maintaining their temperature in more various states of the external atmosphere. Third. Finally, the actions of the cerebro-spinal system, intellectual and muscular, are more regular and powerful when not liable to interruption from the operations of the ganglionic nerves, and the visceral functions presided over by them. When the boa-constrictor digests, he falls into a state of torpor that exceeds in degree, but not in kind, the drowsy rumination of a cow chewing her cud. Such animals are slaves to their nutritive functions, by which those of the brain and spinal cord may at any time be, as it were, oppressed and overwhelmed. The capacity for independence increases with every rise in the hierarchical scale of vertebrates, until it culminates in man—able to think and talk over his dinner; to manufacture heat in his limbs while drawing blood to his cerebral hemispheres; to sustain in complete unconsciousness innumerable delicate and complicated chemical metamorphoses in all the tissues of his body, while concentrating every conscious effort of his mind upon equally delicate processes of thought and will.

The peculiarities that, when coarsely emphasized, serve to distinguish different species of animals from one another, are repeated in more subtle gradations, as varieties among the different classes, and even different individuals of the human race. Here may be found, at least, faint echoes and distant reminiscences of facts that stand out in bold relief throughout the animal kingdom. The classification of sex is certainly one of those that offer an interesting opportunity for such comparison, especially in regard to the relations existing between the operations of the ganglionic, and those of the cerebro-spinal system. As the authors who have asserted the complete subordination of the brain to the instincts in woman, have thus, perhaps unconsciously, reduced her to the anatomical level of the crustacea; so those who, like Dr. Clarke, insist on the incompatibility between cerebral action and the process of ovulation, imply a predominance of ganglionic activity in women that must render them the physiological inferiors of the animals or individuals in whom no such incompatibility exists.

Were such opposition between cerebral and ganglionic functions only noted when a rhythmical intermittence was introduced into the latter, and were such rhythm observed only in the phenomena of menstruation, it might indeed be possible to fix upon women a peculiar mark of physiological inferiority, almost sufficient to amount to a stigma. But rhythmical movement is characteristic of all physiological actions—of the beating of the heart, the secretions of the stomach, the congestion of the spleen, the circulation of the brain, quite as decidedly as of the ripening of cells in the ovary. The tidal waves described by Michelet have become the exclusive theme of his eloquence, mainly because his attention was not attracted to any but those connected with the more obvious phenomena of menstruation. But many tidal waves rise and cross each other in shorter or longer cycles—waves of pulse and of temperature, of sleep and wakefulness, intermittences of secretion and excretion. In regard to the latter, it is noticeable that an intermittent excretion, as of bile or urine, is provided for by a continuous secretion, and that the same is true of the excretion upon whose rhythm an erroneously exceptional emphasis has been laid—that of the menstrual fluid. Here, as elsewhere, the intermittent phenomenon is preceded by long-continued cell growth—effected by precisely such processes of cellular assimilation and metamorphosis as take place in the elements of the liver and the kidneys. The cell growth in question is effected in the ovaries; the final stage of the process, the rupture of the containing cell or ovisac, and escape of the ovule, is attended by a concentration of nervous activity in the ganglionic masses sending nerves to those organs—analogous to that which occurs in the solar plexus at periods of digestion; the fall of the ovule is itself analogous to the shedding of epithelial cells in the gastric follicles; the afflux of blood to the utero-ovarian veins, analogous to the periodical congestion of the gastro-splenic vascular apparatus. Only, in this last case, the congestion results in the elaboration of a fluid secretion, the gastric juice; in the utero-ovarian plexus, where no secretion is required, the blood itself is discharged. It is difficult, with these facts, to understand the assertion that. “Periodicity is the grand (i.e. exclusive) characteristic of the female sex.”

In normal conditions, the process of digestion and of menstruation are both accomplished without invading the consciousness of the individual whose body is the theatre of such extraordinary phenomena. Various abnormal conditions raise the one or the other to the sphere of consciousness—various stages in their evolution. Consciousness of nutritive functions is always painful, and digestion, quite as well as ovulation, may become a process most disturbing to cerebral tranquility and efficiency. The longer duration of the latter is compensated by the more frequent occurrence of the former. The ovaries are decidedly active during at least fifteen days of every month; the stomach, during three or four hours after each meal, or from nine to twelve hours a day. As a matter of fact the digestive function is much more often the occasion of conscious discomfort, than is the function of ovulation. Whenever it becomes so, the dyspeptic approaches the condition of the reptiles or ruminating animals, in whom the process of digestion so absorbs the powers of the nervous system that all other modes of its activity are suspended. But such a condition is universally regarded as an evidence of disease, nor could any considerations concerning the complexity and importance of the ganglionic nerves of the stomach, or the intermittent character of digestion, convert the misfortune of the dyspeptic into a physiological type for the race. At the most may it be admitted:

1st. That in civilized communities dyspepsia is a very common disease.

2d. That dyspeptics require rest of mind and body to facilitate the laborious process of digestion.

Cæteris paribus, these same propositions may be held of those suffering from abnormal modes of activity in another part of the ganglionic system—that connected with menstruation. A third proposition is, moreover, common to both, namely, that repose of the cerebro-spinal system is not required throughout the entire period of ganglionic activity, unless in exceptionally morbid cases. Thus, the process of digestion occupies from three to five hours, but an hour's repose after dinner is generally sufficient to avert discomfort. Similarly, the process of ovulation continues over fifteen days—menstruation lasts from three to six—but even in the cases that demand rest, six to twelve hours is usually enough, and more than enough.

It is noticeable that a slighter disturbance of normal conditions is needed to render digestion painful than to cause painful ovulation, that is, pain preceding the menstrual flow. Pain in menstruation, which is much more frequent, is dependent upon other conditions than the activity of the ovaries, and lasts a very much shorter time than does either the function of ovulation, or even than the uterine congestion secondary to it. Outside of actual uterine disease, the pain at this moment is most often dependent on uterine cramp, itself excited by a spasmodic contraction of blood-vessels that interfere with its circulation. As these remarks are addressed to non-medical readers, a word of explanation is here necessary.

It has been shown by experiment that the sudden arrest of the circulation in muscular fibre is sufficient to induce in the latter violent contractions. Thus, the cramps of the legs in cholera patients are due to the stagnation of blood in their muscles. These cramps are even more easily induced in the muscular fibre of the viscera—the unstriped, involuntary muscles—such as exist in the intestine, bladder, and uterus. Anything that will cause a sudden contraction of the blood-vessels in the uterus will, therefore, by cutting off the supply of blood, cause the muscular fibre of the uterus to contract in painful cramps. The small blood-vessels are themselves provided with circular muscular fibres, whose contraction necessarily draws the walls of the vessels together, obliterates their canal, and shuts out the blood. This contraction is effected by stimulation of the fine nerves, called vaso-motor, that are distributed to these muscular fibres, and which are derived from the sympathetic ganglia, that form part of that same ganglionic system from which the nerves of the ovaries and other viscera are supplied. The utero-ovarian blood-vessels derive their nerves from the hypogastric plexus, which, formed by branches from both sympathetic ganglia and spinal cord, is the exclusive source of the innervation of the uterus and ovaries. The ganglionic nervous excitement coincident with the maturation of the ovule and the congestion of the uterus, is easily communicated to the vaso-motor nerves of the latter organ. At the very moment, therefore, that the uterine blood-vessels are dilated, and blood is being exhaled into the uterine cavity, an excessive stimulation of the vaso-motor nerves may cause the blood-vessels to contract; the flow is then temporarily arrested, the circulation in the uterus disturbed, and its muscular fibres thrown into cramps.