Fig. 29.—Functional areas of the cerebral cortex, left hemisphere. (A. A. Stevens.)
The brain is the seat of intelligence and will, the center of all voluntary action. Molecular change in some part of the cerebral substance is the indispensable accompaniment of every phenomenon of consciousness. Indeed, the brain is never in a state of complete repose, there being dreams even during sleep. The brain is not sensitive to injury in the sense of pain. It can be lacerated without much pain.
Various centers exist in the brain, of which the most important perhaps is the motor center. The visual center is in the occipital lobe, the auditory center in the temporal lobe, the [speech center] in the third left frontal convolution. Thus the impulses of the senses have been located, though the function of many parts, the so called silent areas, are still in obscurity.
The [motor center], that is, the center for motion of the skeletal muscles, is situated about the fissure of Rolando and is divided into three parts, one for the legs, one for the face, and one for the arms, the one for the legs being uppermost and the others below in the order mentioned. Fibers from these cells extend down through the brain and cord to the muscles, the fibers being collected into well-recognized bundles and the whole known as the motor tract. There may be one long fiber from a cell in the brain down through most of the cord or there may be a succession of shorter fibers that are not actually connected but are in close contact with each other. In the upper pons the fibers for the face cross to the opposite side, while the rest keep on down through the medulla, and as they emerge from the medulla they too cross to the other side and keep on down in the crossed pyramidal tract. A few fibers do not cross but come down the direct pyramidal tract, which, however, disappears part way down. The crossed pyramidal tract is the true motor tract and in it the fibers are continually sending branches to the cells in the gray matter, where they connect with the anterior horn.
The anatomy of the sensory tract is not so well understood. By it impulses are sent to the brain by the peripheral organs, practically the surface of the body. The sensory fibers connect with the sensory cells in the posterior horn, from which fibers are sent to the brain, practically the reverse of motor action. There are three chief sensory tracts, which are supposed to transmit different sensations, one pain, one muscular sensations, and the third sensations of touch. All these tracts, of which the chief is the direct cerebellar tract, in passing up the cord pass to the opposite side at different levels and then go on to the cortex of the brain.
The action of the nerves is similar to reflex action, only that an effort of will is needed to send an impulse from the brain. It is by the help of the brain along this line that an infinity of artificial reflexes or habits is acquired, for which volition is needed in the beginning but which are later done unconsciously. Herein lie the possibilities of all education.
The brain and spinal cord work together, the cord acting as a medium between the brain, in which all the higher psychical processes, such as will, thought, etc., originate, and the muscular apparatus. The cord, however, has some action entirely independent of the brain, as is seen in reflex action. This action is entirely involuntary, so that the cord is sometimes spoken of as the seat of involuntary action, commonly called reflex action. All unconscious acts are reflex acts, as when the hand is drawn away from a hot iron. If an impulse is sent along one of the sensory fibers, it enters the cord through the posterior horn, where its nerve cell is found. Then, through some connection between the nerve cell of the sensory fiber and that of the motor fiber the impulse is transmitted to the motor cell and another impulse is sent out of the cord along the motor fiber of the nerve to the muscle. One of the commonest reflexes is the knee-jerk. Reflex action is important because the reflexes are interfered with, delayed, destroyed, or increased in different diseases. The time normally required for a reflex act is very brief, that for the knee-jerk being about three one-hundredths of a second.
The nerves of the head, known as the cranial nerves, arise from the brain, while the rest of the body is supplied by the spinal nerves, which come off at intervals from the spinal cord. The cranial nerves consist of twelve pairs: (1) The olfactory or nerve of smell, (2) the optic or nerve of sight, (3) the motor oculi, (4) the patheticus, which controls the eye, (5) the trigeminus or trifacial, a nerve of general sensation, motion, and taste, (6) the abducens, a motor nerve, (7) the facial nerve of the face, ear, palate, and tongue, (8) the auditory or nerve of hearing, (9) the glosso-pharyngeal, nerve of sensation and taste, (10) the pneumogastric or vagus, which is both motor and sensory and governs respiration, the heart, and the stomach, (11) the spinal accessory, to the muscles of the soft palate, and (12) the hypoglossal, the motor nerve to the tongue.
The spinal nerves also are arranged in pairs: Eight cervical pairs, twelve dorsal or thoracic, five lumbar, five sacral, and one coccygeal, these titles denoting their point of origin near the vertebra of the same name. Each of these nerves arises by two roots, an anterior motor root from the anterior horn of gray matter and a posterior sensory root from the posterior horn, the latter having a ganglion upon it. After emerging from the cord the two roots unite to form the nerve, that the nerve may contain both motor and sensory fibers. The motor fibers are called efferent because they carry impulses from the cord, while the sensory are called afferent because they carry impulses back to the cord. After leaving the cord the nerves unite to form plexuses, which again divide into various nerve trunks and are distributed to the muscles.
The first cervical nerves pass out of the spinal column above the first cervical vertebra and the other cervical nerves below that and the succeeding vertebræ, while the other spinal nerves emerge each below the corresponding vertebra, as the first dorsal below the first dorsal vertebra, etc. After emerging they break up into a large anterior division and a small posterior division, the posterior branches supplying the spine and the dorsal muscles and skin, the anterior the rest of the trunk and the limbs. The cervical plexus is formed by the anterior divisions of the first four cervical nerves, the brachial plexus by the last four cervical and the first dorsal or thoracic nerves, the lumbar plexus by the four upper lumbar, and the sacral plexus by the last lumbar and the four upper sacral nerves.