In the orange-pulp these cells contain only sweet juice, but in other parts of the orange-tree or any other plant they contain a sticky substance with little grains in it. This substance is called "protoplasm," or the first form of life, for it is alive and active, and under a microscope you may see in a living plant streams of the little grains moving about in the cells.
Now we are prepared to explain how our plant grows. Imagine the tiny primrose plantlet to be made up of cells filled with active living protoplasm, which drinks in starch and other food from the seed-leaves. In this way each cell will grow too full for its skin, and then the protoplasm divides into two parts and builds up a wall between them, and so one cell becomes two. Each of these two cells again breaks up into two more, and so the plant grows larger and larger, till by the time it has used up all the food in the seed-leaves, it has sent roots covered with fine hairs downwards into the earth, and a shoot with beginnings of leaves up into the air.
Sometimes the seed-leaves themselves come above the ground, as in the mustard-plant, and sometimes they are left empty behind, while the plantlet shoots through them.
And now the plant can no longer afford to be idle and live on prepared food. It must work for itself. Until now it has been taking in the same kind of food that you and I do; for we too find many seeds very pleasant to eat and useful to nourish us. But now this store is exhausted. Upon what then is the plant to live? It is cleverer than we are in this, for while we cannot live unless we have food which has once been alive, plants can feed upon gases and water and mineral matter only. Think over the substances you can eat or drink, and you will find they are nearly all made of things which have been alive: meat, vegetables, bread, beer, wine, milk; all these are made from living matter, and though you do take in such things as water and salt, and even iron and phosphorus, these would be quite useless if you did not eat and drink prepared food which your body can work into living matter.
But the plant as soon as it has roots and leaves begins to make living matter out of matter that has never been alive. Through all the little hairs of its roots it sucks in water, and in this water are dissolved more or less of the salts of ammonia, phosphorus, sulphur, iron, lime, magnesia, and even silica, or flint. In all kinds of earth there is some iron, and we shall see presently that this is very important to the plant.
Suppose, then, that our primrose has begun to drink in water at its roots. How is it to get this water up into the stem and leaves, seeing that the whole plant is made of closed bags or cells? It does it in a very curious way, which you can prove for yourselves. Whenever two fluids, one thicker than the other, such as treacle and water for example, are only separated by a skin or any porous substance, they will always mix, the thinner one oozing through the skin into the thicker one. If you tie a piece of bladder over a glass tube, fill the tube half-full of treacle, and then let the covered end rest in a bottle of water, in a few hours the water will get in to the treacle and the mixture will rise up in the tube till it flows over the top. Now, the saps and juices of plants are thicker than water, so, directly the water enters the cells at the root it oozes up into the cells above, and mixes with the sap. Then the matter in those cells becomes thinner than in the cells above, so it too oozes up, and in this way cell by cell the water is pumped up into the leaves.
When it gets there it finds our old friends the sun-beams hard at work. If you have ever tried to grow a plant in a cellar, you will know that in the dark its leaves remain white and sickly. It is only in the sunlight that a beautiful delicate green tint is given to them, and you will remember from Lecture II. that this green tint shows that the leaf has used all the sun-waves except those which make you see green; but why should it do this only when it has grown up in the sunshine?
The reason is this: when the sunbeam darts into the leaf and sets all its particles quivering, it divides the protoplasm into two kinds, collected into different cells. One of these remains white, but the other kind, near the surface, is altered by the sunlight and by the help of the iron brought in by the water. This particular kind of protoplasm, which is called "chlorophyll," will have nothing to do with the green waves and throws them back, so that every little grain of this protoplasm looks green and gives the leaf its green colour.
It is these little green cells that by the help of the sun-waves digest the food of the plant and turn the water and gases into useful sap and juices. We saw in Lecture III. that when we breathe-in air, we use up the oxygen in it and send back out of our mouths carbonic acid, which is a gas made of oxygen and carbon.
Now, every living things wants carbon to feed upon, but plants cannot take it in by itself, because carbon is solid (the blacklead in your pencils is pure carbon), and a plant cannot eat, it can only drink-in fluids and gases. Here the little green cells help it out of its difficulty. They take in or absorb out of the air carbonic acid gas which we have given out of our mouths and then by the help of the sun-waves they tear the carbon and oxygen apart. Most of the oxygen they throw back into the air for us to use, but the carbon they keep.