395. Moreover, although observation may show that a seedling, weighing only two or three grains, may double its bulk and weight every week of its early growth, and may in time produce a huge amount of vegetable matter, it is still to be asked what this vegetable matter is, where it came from, and by what means plants are able to increase and accumulate it, and build it up into the fabric of herbs and shrubs and lofty trees.
[396.] Protoplasm. All this fabric was built up under life, but only a small portion of it is at any one time alive. As growth proceeds, life is passed on from the old to the new parts, much as it has passed on from parent to offspring, from generation to generation in unbroken continuity. Protoplasm is the common name of that plant-stuff in which life essentially resides. All growth depends upon it; for it has the peculiar power of growing and multiplying and building up a living structure,—the animal no less than the vegetable structure, for it is essentially the same in both. Indeed, all the animal protoplasm comes primarily from the vegetable, which has the prerogative of producing it; and the protoplasm of plants furnishes all that portion of the food of animals which forms their flesh and living fabric.
397. The very simplest plants (if such may specifically be called plants rather than animals, or one may say, the simplest living things) are mere particles, or pellets, or threads, or even indefinite masses of protoplasm of vague form, which possess powers of motion or of changing their shape, of imbibing water, air, and even other matters, and of assimilating these into plant-stuff for their own growth and multiplication. Their growth is increase in substance by incorporation of that which they take in and assimilate. Their multiplication is by spontaneous division of their substance or body into two or more, each capable of continuing the process.
398. The embryo of a phanerogamous plant at its beginning ([344]) is essentially such a globule of protoplasm, which soon constricts itself into two and more such globules, which hold together inseparably in a row; then the last of the row divides without separation in the two other planes, to form a compound mass, each grain or globule of which goes on to double itself as it grows; and the definite shaping of this still increasing mass builds up the embryo into its form.
Fig. 433-436. Figures to illustrate the earlier stages in the formation of an embryo; a single mass of protoplasm (Fig. 433) dividing into two, three, and then into more incipient cells, which by continued multiplication build up an embryo.
[399.] Cell-walls. While this growth was going on, each grain of the forming structure formed and clothed itself with a coat, thin and transparent, of something different from protoplasm,—something which hardly and only transiently, if at all, partakes of the life and action. The protoplasm forms the living organism; the coat is a kind of protective covering or shell. The protoplasm, like the flesh of animals which it gives rise to, is composed of four chemical elements: Carbon, Hydrogen, Oxygen, and Nitrogen. The coating is of the nature of wood (is, indeed, that which makes wood), and has only the three elements, Carbon, Hydrogen, and Oxygen, in its composition.