The fourth and shortest essay on the “Transformation of the Axolotl into Amblystoma,” starts primarily with the intention of showing that cases of sudden transformation are no proof of per saltum development. When this essay first appeared the view was still widely entertained that we had here a case proving per saltum development. That this explanation was erroneous is now generally admitted, but I believe that those who suppose that we have here to deal with some quite ordinary phenomenon which requires no explanation, now go too far towards the other extreme. The term “larval reproduction” is an expression, but no explanation; we have therefore to attempt to find out the true interpretation, but whether the one which I have given is correct must be judged of by others.

These four essays lead up to a fifth and concluding one “On the Mechanical Conception of Nature.” Whilst the results obtained are here summed up, it is attempted to form them into a philosophical conception of Nature and of the Universe. It will be thought by many that this should have been left to professed philosophers, and I readily admit that I made this attempt with some misgiving. Two considerations, however, induced me to express here my own views. The first was that the facts of science are frequently misunderstood, or at any rate not estimated at their true value, by philosophers;[2] the second consideration was, that even certain naturalists and certainly very many non-naturalists, turn distrustfully from the results of science, because they fear that these would infallibly lead to a view of the Universe which is to them unacceptable, viz. the materialistic view. With regard to the former I wished to show that the views of the development of organic Nature inaugurated by Darwin and defended in this work are certainly correctly designated mechanical; with reference to the latter I wished to prove that such a mechanical conception of the organic world and of Nature in general, by no means leads merely to one single philosophical conception of Nature, viz. to Materialism, but that on the contrary it rather admits of legitimate development in a quite different manner.

Thus in these last four essays much that appears heterogeneous will be found in close association, viz. scientific details and general philosophical ideas. In truth, however, these are most intimately connected, and the one cannot dispense with the other. As the detailed investigations of the three essays find their highest value in the general considerations of the fourth, and were indeed only possible by constantly keeping this end in view, so the general conclusions could only grow out of the results of the special investigations as out of a solid foundation. Had the new materials here brought together been already known, the reader would certainly have been spared the trouble of going into the details of special scientific research. But as matters stood it was indispensable that the facts should be examined into and established even down to the most trifling details. The essay “On the Origin of the Markings of Caterpillars” especially, had obviously to commence with the sifting and compilation of extensive morphological materials.

August Weismann.

Freiburg in Baden,
November, 1881.


CONTENTS.

Part I.

ON THE SEASONAL DIMORPHISM OF BUTTERFLIES.