This shows the capital importance of conjugation and of the substance of the conjugated nuclei, especially of their chromatin. The fact that, in certain of the lower animals, the protoplasm of the egg without nuclei may occasionally produce some phenomena of cell division, thanks to its inherited mnemic engrams, in no way alters the fundamental principle which alone occurs in man, for this vicarious action, which is moreover rudimentary, only happens when the protoplasm of the egg is not consumed by the conjugated nuclei.
Parthenogenesis is also a very interesting phenomenon in the history of our animal ancestors, but for the same reasons it has no direct interest for humanity.
If we take into consideration all the observations of which we have just spoken, which are as simple as they are irrefutably demonstrated, it is hardly possible to interpret them in any other way than by the following hypothesis:
In each sexual gland, male or female, the germinal cells which are produced by division of the cells of the embryo, reserved primarily for reproduction, differ considerably from each other in quality and contain in their infinitely small atoms very diverse and irregularly distributed energies, inherited from their different ancestors. Some contain more paternal and others more maternal energy, and among the former there are some contain, for example, more paternal grandfather and others more maternal grandmother, and so on to infinity, till it is impossible to discover the ancestral origin of the fully grown individual we are examining. The same holds good for the energies of the maternal cells.
At the time of conjugation, the qualities of the child which will result from it depend therefore on conditions of the ancestral qualities of the conjugated egg and spermatozoön. Moreover, although of the same size, the nuclei which become conjugated are evidently of unequal strength; the energies of one or the other predominate later on in the embryo, and still later in man. According to circumstances the latter will resemble more or less his paternal or maternal progenitors.
Moreover, the different organs of the body may receive their energies from different parts of the conjugated nuclei in different degrees. A person may have his father's nose and his mother's eyes, the paternal grandmother's humor and the maternal grandfather's intelligence, and all this with infinite degrees and variations, for it is only a matter of more or less accentuated averages. In my own face the two halves are distinctly different, one resembling my maternal ancestry and the other, in a lesser degree, my paternal ancestry, these points being seen distinctly in photographs taken in profile.
Each germinal cell contains the hereditary mneme of its ancestors, paternal and maternal, and the two cells united by conjugation (Fig. 17) that of the ancestors of each of them. We have spoken above of ecphorias produced according to Mendel's law and reproducing characters which have been latent during one or two generations. Darwin was the first to study this interesting fact, which shows how atavism often results from the crossing of varieties. There are several varieties of fowls which do not brood; if two of these varieties, b and c, are crossed excellent brooders are obtained. Semon assumes that in each of the non-brooding varieties the ancestral energy, a, of the primary species, is weaker than that of varieties b and c; we have then a > b, and a < c. But if b is coupled with a the product represents the value b + c + a + a. Then b and c are in equilibrium; and a being doubled becomes stronger than each of them and arrives at ecphoria in their place, which restores the faculty of brooding to the product of crossing.
De Vries has shown, in the crossing of varieties with their primary species, more or less analogous phenomena which he calls "Vicino-variations." Conjugation leads to infinite combinations and variations which the law of heredity traverses like a guiding line.
The celebrated zoölogist, Weismann, considers that the chromatin of each germinal cell contains a considerable quantity of particles each of which is capable of forming an entire organism similar to the parents; these he calls "ides." According to Weismann, each ide is subdivided into "determinants" from which each part of the body is derived, being potentially predetermined in them. According to the action of a yet unknown irritation male or female determinants develop in each individual of the animal species with separate sexes. But if the determinants are disordered, either by abnormal variations or by pathological causes, hermaphrodites or monstrosities may be produced. In animals which are normally hermaphrodite (snails, etc.), there is only one kind of sexual determinant, while in polymorphous animals (ants, etc.), there are as many as the polymorphous forms. The conception of "ides" and "determinants" is only a hypothesis to which we must not attach much value. The mnemic laws established by Semon give a much better explanation of the facts.
It has often been maintained that the qualities of higher forms of man are exhausted in a few generations, while the mass of mediocrities continually produce new genius. The fact that the descendants of distinguished men are often mediocre and that remarkable men suddenly arise from the common people, appears at first sight to support this superficial assertion. It is forgotten, however, that in a people whose average mass consists of thousands or millions of individuals, while men of higher powers are only counted by units or dozens, all this arithmetic is reduced to absurdity by the inequality of numbers, as soon as the law of heredity is understood. To make a more exact calculation, it would be necessary to compare the number of superior men who have arisen from some hundreds of the most distinguished families of a country with that of distinguished men who have arisen from some millions of the rest of the people, and then calculate the percentage. It is also necessary to take into account the means employed in the education of the individuals. If education is obligatory and gratuitous in a country, this factor will have less importance.