Adaptation must be studied, not only with reference to temperature ranges and rainfall, but especially with reference to atmospheric humidity and requirements of transpiration.

Many species have a wide range of natural distribution, and hence of climatic adaptation. If such are to be used, it is important to secure seeds from that part of the range of natural distribution where the plants must be hardiest, i. e., the coldest and driest region in which it occurs, which insures hardy qualities in the offspring. For instance, the Douglas spruce from the humid and evenly tempered Pacific Slope will not be as hardy as that grown from seed collected on the dry and frigid slopes of the Rockies. Lack of attention to this requisite accounts for many failures. It must also be kept in mind that, while a species may be able to grow in another than its native climate, its wood may not there have the same valuable qualities which it develops in its native habitat.

Adaptability to soil must be studied less with reference to mineral constituents than to physical condition. Depth and moisture conditions, and the structure of the soil, which influences the movement of water in it, are the most important elements. While all trees thrive best in a moist to "fresh" soil of moderate depth (from 2 to 4 feet) and granular structure, some can adapt themselves to drier or wetter, shallow, and compact soils. Fissures in rocks into which the roots can penetrate often stand for depth of soil, and usually aid in maintaining favorable moisture conditions. In soils of great depth (i. e., from the surface to the impenetrable subsoil) and of coarse structure water may drain away so fast as not to be available to the roots.

Soil moisture must always be studied in conjunction with atmospheric moisture; for, while a species may thrive in an arid soil, when the demands of transpiration are not great, it may not do so when aridity of atmosphere is added. Trees of the swamp are apt to be indifferent to soil moisture and to thrive quite well, if not better, in drier soils.

Adaptability to site.—While a species may be well adapted to the general climatic conditions of a region, and in general to the soil, there still remains to be considered its adaptability to the particular "site," under which term we may comprise the total effect of general climate, local climate, and soil. The general climatic conditions are locally influenced, especially by the slope, exposure, or aspect, and the surroundings. Thus we know that eastern exposures are more liable to frost, western exposures more liable to damage from winds, southern more apt to be hot and to dry out, and northern to be cooler and damper, having in consequence a shorter period of vegetation. Hollows and lowlands are more exposed to frosts and more subject to variations in soil moisture, etc.

Hence for these various situations it is advisable to select species which can best withstand such local dangers.

The use value, or utility, of the species is next to be considered. This must be done with reference to the commercial and domestic demand, and the length of time it takes the species to attain its value. The greater variety of purposes a wood may serve—i. e., the greater its general utility—and the sooner it attains its use value the better. White pine for the northeastern States as a wood is like the apple among fruits, making an all-round useful material in large quantities per acre in short time. Tulip poplar, applicable to a wider climatic range, is almost as valuable, while oak, ash, and hickory are standard woods in the market. Other woods are of limited application. Thus the black locust, which grows most quickly into useful posts, has only a limited market, much more limited than it should have; hickory soon furnishes valuable hoop poles from the thinnings, and later the best wagon material, not, however, large quantities in a short time; while black walnut of good quality is very high in price, the market is also limited, and the dark color of the heartwood, for which it is prized, is attained only by old trees. The black cherry, used for similar purposes, attains its value much sooner.

By planting various species together, variety of usefulness may be secured and the certainty of a market increased.

The forest value of the species is only in part expressed by its use value. As has been shown in another place, the composition of the crop must be such as to insure maintenance of favorable soil conditions, as well as satisfactory development of the crop itself. Some species, although of high use value, like ash, oak, etc, are poor preservers of soil conditions, allowing grass and weeds to enter the plantation and to deteriorate the soil under their thin foliage. Others, like beech, sugar maple, box elder, etc., although of less use value, being dense foliaged and preserving a shady crown for a long time, are of great forest value as soil improvers.