Essay on the Theory of the Earth

by baron Georges Cuvier

First, as to size, the difference is very remarkable, it not being uncommon to find the fossil horns ten feet between the extreme tips[423], while the largest elk’s horns never measure four feet. This measurement in a pair in the Museum of the Royal Dublin Society, is three feet seven inches: the largest pair seen by Pennant in the house of the Hudson’s Bay Company, measured thirty-four inches[424].

The horn of the elk has two palms, a lesser one which grows forward from the front of the beam, where the principal palm begins to expand. This is called brow antler by Cuvier, but it corresponds in situation rather to the sur-antler, there being, properly speaking, no brow antler attached to the root of the beam. The elk has no posterior antler similar to that of the fossil animal, nor does its beam take a similar arched direction, but runs more directly outwards.

Cuvier remarks, that the palm of the fossil horn increases in breadth as it extends outwardly, while that of the elk is broadest next the beam.

The palm of the elk’s horn is directed more backwards, while the fossil one extends more in the lateral direction. The antlers of the elk are shorter and more numerous than those of the fossil animals.

As the horns of the fossil animal exceed in size those of the elk, so, on the contrary, does the skull of the latter exceed in size that of the former; the largest heads of the fossil species not exceeding one foot nine inches in length, while the head of the elk is frequently two feet. The fossil head is broader in proportion; its length being to its breadth as two to one; in the elk they are as three to one, according to Parkinson.[425] The breadth of the skull between the roots of the horns is but four inches in the fossil skulls; in that of the elk in the Society’s Museum it is 6½ inches.

Cuvier thinks it probable that the females of the fossil species had horns[426], an opinion to which I am very much disposed to subscribe, from having observed that these parts present differences in size and strength, which appear not to be dependent on differences of age. For instance, the teeth of the specimen in Trinity College are much more worn down, and the sutures of the skull are more effaced than in the specimen described in this paper; yet the horns of the latter are much more concave, and more expanded, than those of the former; and on comparing a single horn of each of these specimens together, that belonging to the Society exceeds the other by nearly a sixth in the length, and little less than a third in the breadth; it is not, therefore, unlikely that the animal whose horns were larger and more curved was a male. Something similar to this is observed in the rein-deer, both sexes of which have horns, but with this difference, that they are smaller and less branched in the female. Hence we find that this animal possessed characters of its own sufficient to prove it of a species as distinct from the moose or elk as this latter species is from the rein-deer or any other. Therefore, it is improper to retain the name of elk or moose deer any longer: perhaps it might be better called the Cervus megaceros, a name merely expressive of the great size of its horns.

That this animal shed its head furniture periodically, is proved by the occasional occurrence of detached horns having the smooth convex surface below the burr, similar to what is observed on the cast horns of all deer. Specimens of this are to be seen in the Museum of Trinity College, and I possess one myself, of which I have had a drawing made. As every other species of deer shed their horns annually, there is no reason for supposing that that process occurred at longer intervals in this.

It is a popular opinion with the Indians that the elk is subject to epilepsy, with which he is frequently seized when pursued, and thus rendered an easy prey to the hunters. Many naturalists affect to disbelieve this account, without, however, assigning any sufficient reason. But if it be considered, that, during the growth of the horns, there must be a great increased determination of blood to those parts, which are supplied by the frontal artery, a branch from the internal carotid, it is quite conformable to well established pathological principles, to suppose, that, after the horns are perfected, and have ceased to receive any more blood, that fluid may be determined to those internal branches of the carotid which supply the brain, and establish a predisposition to such derangements of its circulation as would produce epilepsy, or even apoplexy: if such an effect were produced in consequence of the size of the horns in the elk, it is reasonable to suppose that it prevailed in a greater degree in the fossil animal whose horns were so much larger.

What could have been the use of these immense horns? It is quite evident that they would prevent the animal making any progress through a thickly wooded country, and that the long, tapering, pointed antlers were totally unfit for lopping off the branches of trees, a use to which the elk sometimes applies his horns[427], and for which they seem well calculated, by having their antlers short and strong, and set along the edge of the palm, somewhat resembling the teeth of a saw in their arrangement. It would rather appear, then, that they were given the animal as weapons for its protection, a purpose for which they seem to have been admirably designed; for their lateral expansion is such, that should occasion require the animal to use them in his defence, their extreme tips would easily reach beyond the remotest parts of his body; and if we consider the powerful muscles for moving the head, whose attachments occupied the extensive surfaces of the cervical vertebræ, with the length of the lever afforded by the horns themselves, we can easily conceive how he could wield them with a force and velocity which would deal destruction to any enemy having the hardihood to venture within their range.