From G. bursarius industrius, major differs in color, being Prout's Brown, instead of Cinnamon Brown (less Fuscous); body averaging 10 per cent longer; total length in males from 9 to 9.7 per cent longer, hind foot 9.7 per cent longer on the average; skull averaging larger in all parts measured. Occiput less inclined anterodorsally; top nearly flat, less arched than that of G. b. industrius; auditory bulla averaging slightly larger and less inflated.
Remarks.—Specimens of this subspecies from Norman, Cleveland County, Oklahoma, and Canton, Dewey County, Oklahoma, and most of those from Kansas, are more Fuscous than topotypes and tend toward G. bursarius majusculus. Specimens from McPherson County have a darker dorsal stripe resembling that of G. bursarius majusculus. One adult from Little Salt Marsh, Stafford County, is pale, closely resembling topotypes.
Most of the cranial characters, nevertheless, are constant in all available specimens, except that in specimens of each sex from the type locality the basilar length averages 4 to 5 per cent shorter. In the constancy of size of the relatively large auditory bullae and in the nearly flat dorsal profile of the cranial part of the skull, the specimens from Kansas agree with the specimens from the type locality.
Specimens from Harper County have the occiput slightly inclined anterodorsally and thus are reminiscent of industrius which has an even greater inclination of the occiput. Probably the appearance in dilute fashion of this character in Harper County is properly to be interpreted as intergradation with industrius. If so, the actual intergradation may be to the northwest via Pratt County since specimens from Barber County, immediately west of Harper and lying between Harper County and the range of industrius, do not have the occiput so inclined.
Of a pair of adults from eight miles west of Rosalia, Butler County, the female is indistinguishable in color from adults of G. b. industrius from northern Meade County and from two specimens from eleven miles west of Clarendon, Donley County, Texas, near the type locality of G. b. major. The male from eight miles west of Rosalia is darker as compared either with G. b. industrius or G. b. major and the coloration of the upper parts resembles those in G. b. majusculus; the underparts are only slightly paler than the upper parts as in majusculus. Measurements of the skulls are intermediate between the averages for G. b. majusculus and those for G. b. major. These specimens from eight miles west of Rosalia are intermediate structurally, and since they are intermediate geographically between G. b. majusculus and G. b. major, they suggest intergradation of the two subspecies. The specimens in question are referred to major because the size is nearer that of major. It is mainly the intermediate nature of these two specimens from Butler County, and the intermediate nature of the specimens from McPherson County, Kansas, that have caused us to treat G. b. majusculus as only subspecifically distinct from the more western subspecies, major.
Specimens examined.—Total number, 77, as follows: Ellsworth County: 2 mi. S Ellsworth, 1. McPherson County: Smoky Hill River, 1 mi. S and 1/2 mi. W Lindsborg, 5; 1/2 mi. E McPherson, 1. Stafford County: Little Salt Marsh, 12; no locality more precise than county, 3. Reno County: 8 mi. N and 1 mi. E Haven, 2. Harvey County: 1 mi. E and 1/2 mi. N Halstead, 1; Halstead, 3. Butler County: 8 mi. W Rosalia, 2. Barber County: near South Bridge, Sun City, 1; 2 mi. S Sun City, 1; Wells Ranch, Aetna, 5; "1 mi. W Aetna," 3; near South Bridge, Aetna, 1; near Bridge, 1 mi. S Aetna, 2. Harper County: 4-1/2 mi. NE Danville, 8; 1 mi. N Harper, 11; 3 mi. S Harper, 1. Cowley County: 3 mi. SW Arkansas City, 4; 3 mi. SE Arkansas City, 9; 3 mi. S Arkansas City, 1.
Measurements of Adult Males of Geomys
(In millimeters)
| Key for table headings in table on this page. | |||||||||||||
| N: Number of individuals averaged or catalogue number | |||||||||||||
| L: Total length | |||||||||||||
| T: Length of tail | |||||||||||||
| H: Length of hind foot | |||||||||||||
| B: Basilar length | |||||||||||||
| Na: Length of nasals | |||||||||||||
| Z: Zygomatic breadth | |||||||||||||
| M: Mastoid breadth | |||||||||||||
| Rb: Breadth of rostrum | |||||||||||||
| I: Interorbital constriction | |||||||||||||
| A: Alveolar length of maxillary tooth row | |||||||||||||
| E: Extension of premaxilla posterior to nasals | |||||||||||||
| S: Depth of skull | |||||||||||||
| Rl: Length of rostrum | |||||||||||||
| N | L | T | H | B | Na | Z | M | Rb | I | A | E | S | Rl |
G. b. lutescens; topotypes | |||||||||||||
| 5 ave. | 266 | 82.0 | 34.2 | 40.0 | 17.7 | 30.5 | 26.8 | 11.5 | 6.7 | 8.6 | 3.9 | 17.1 | 20.8 |
| min. | 257 | 76.0 | 33.0 | 38.3 | 16.0 | 29.1 | 26.1 | 11.2 | 6.3 | 8.1 | 3.5 | 16.2 | 19.1 |
| max. | 276 | 91.0 | 36.0 | 42.4 | 20.3 | 31.7 | 27.5 | 11.9 | 6.9 | 9.2 | 4.2 | 17.7 | 23.6 |
2 mi. NE Ludell, Rawlins Co., Kansas | |||||||||||||
| 12088 | 272 | 92.0 | 35.0 | 43.2 | 19.1 | 32.3 | 27.7 | 11.3 | 6.6 | 8.4 | 2.8 | 18.0 | 22.1 |
G. b. majusculus; Douglas Co., Kansas | |||||||||||||
| 16 ave. | 289 | 79.8 | 36.3 | 47.1 | 21.0 | 34.1[A] | 30.4 | 12.1 | 6.8 | 9.3 | 3.7 | 18.5 | 24.9 |
| min. | 273 | 70.0 | 32.0 | 44.7 | 18.9 | 30.5 | 27.5 | 11.1 | 6.5 | 8.2 | 2.9 | 17.3 | 22.9 |
| max. | 308 | 95.0 | 55.0 | 49.9 | 23.2 | 38.0 | 34.5 | 13.5 | 7.6 | 10.3 | 5.7 | 20.0 | 28.1 |
G. b. jugossicularis; Morton Co., Kansas | |||||||||||||
| 4 ave. | 265 | 82.0 | 34.2 | 40.7 | 16.9 | 30.0 | 27.9 | 10.7 | 6.0 | 8.6 | 5.2 | 17.3 | 21.2 |
| min. | 250 | 68.0 | 30.0 | 38.5 | 16.1 | 29.0 | 27.5 | 10.5 | 5.5 | 8.2 | 4.7 | 16.4 | 20.2 |
| max. | 285 | 92.0 | 37.0 | 42.4 | 17.4 | 31.1 | 28.4 | 11.0 | 6.2 | 9.2 | 5.5 | 17.9 | 22.0 |
G. b. industrius; Meade Co., Kansas | |||||||||||||
| 8 ave. | 265 | 82.0 | 35.0 | 40.9 | 18.1 | 30.0 | 28.0 | 11.0 | 6.2 | 8.8 | 4.3 | 17.7 | 21.8 |
| min. | 247 | 70.0 | 33.0 | 37.9 | 15.5 | 28.2 | 26.5 | 9.9 | 5.7 | 8.0 | 2.9 | 16.8 | 19.5 |
| max. | 280 | 90.0 | 36.0 | 43.4 | 21.0 | 32.4 | 29.5 | 11.6 | 7.0 | 9.1 | 5.2 | 19.1 | 24.2 |
G. b. major; Wells Ranch, Aetna, Barber Co., Kansas | |||||||||||||
| 11724 | 256 | 66.0 | 34.0 | 41.0 | 18.3 | 31.6 | 28.2 | 10.6 | 6.1 | 9.0 | 4.0 | 17.0 | 21.3 |
1 mi. W Aetna, Barber Co., Kansas | |||||||||||||
| 11153 | 240 | 75.0 | 32.0 | 36.7 | 15.7 | 26.9 | 24.6 | 9.9 | 5.9 | 8.8 | 4.0 | 15.0 | 19.5 |
| 11152 | 240 | 65.0 | 32.0 | 36.0 | 14.2 | 26.1 | 25.4 | 10.9 | 5.6 | 8.5 | 5.0 | 15.5 | 18.5 |
3 mi. SE Arkansas City, Cowley Co., Kansas | |||||||||||||
| 12870 | 246 | 76.0 | 32.0 | 42.1 | 16.0[E] | 33.7 | 29.7 | 11.5 | 6.3 | 9.4 | 4.5 | 17.6 | 21.3 |
3 mi. SW Arkansas City, Cowley Co., Kansas | |||||||||||||
| 12892 | 282 | 84.0 | 33.0 | 41.7 | 17.3 | .... | 27.7 | 10.8 | 6.4 | 8.9 | 4.2 | 17.2 | 21.5 |