Skull: Averaging larger in all parts measured, except that premaxillae do not extend so far posteriorly to nasals in either males or females; interorbital constriction slightly narrower in adult females; temporal ridges forming a more prominent sagittal crest in adult males (sagittal crest barely present in some adult males of major from Harper County).
Remarks.—In employing the subspecific name majusculus we are following Swenk (1939:6) who on the basis of larger size differentiated the animals from southeastern South Dakota, the eastern parts of Nebraska and Kansas, and the western and southern parts of Iowa, from G. bursarius bursarius to which he assigned a more northern geographic range. In the absence of comparative materials of the northern subspecies we cannot make an independent decision on the validity of majusculus and recognize that if it is inseparable from G. b. bursarius the latter name will apply to specimens from northeastern Kansas. We are the more uncertain about applying the name majusculus to specimens from eastern Kansas because they average smaller than topotypes. Only at the northeasternmost locality in Kansas (3 mi. N Cummings, Atchison County) do specimens average as large as topotypes of majusculus. Farther southward they become progressively smaller in eastern Kansas, and we interpret this as intergradation with the still smaller subspecies major, to the southwest. The average external measurements of two adult males from Atchison County are: 321-99-35. Thirty-six miles farther south, in Douglas County, 16 adult males average 289-80-36. From Hamilton, Greenwood County, 80 miles farther southwest, nine adult males average 284-83-35. The maximum total length recorded at these three localities is: Atchison County, 342 (1 of 2 specimens), Douglas County, 308 (1 of 16 specimens), Greenwood County, 357 (in coll. of Dr. Glenn C. Rinker and 1 of 15 males of all ages involved). It will be seen, therefore, that although there is a trend to smaller average size toward the southward, the maximum of 357 millimeters total length at Hamilton exceeds the maximum of 352 millimeters recorded by Swenk (1939:3) among 86 males at Lincoln where the recorded average is largest.
Four specimens from Salina (Debold Farm) are intermediate structurally, as they are also geographically, between G. b. majusculus on the one hand and Geomys bursarius lutescens and Geomys bursarius major on the other hand. In color they agree with majusculus, as they do also in width of nasals posteriorly, in more obtuse angle of the rostrum and maxillary arm of the zygomatic arch. They agree with G. b. lutescens in having the occiput inclined anterodorsally, and are intermediate between majusculus and lutescens, but nearer the latter in size of skull and in length of the rostrum relative to its width.
Specimens examined.—Total number, 148, as follows: Clay County: 6 mi. SW Clay Center, 3. Jackson County: 10-1/2 mi. WSW Holton, 1; no locality more precise than county, 1. Atchison County: 3 mi. N Cummings, 2. Jefferson County: Oskaloosa, 1. Leavenworth County: Fort Leavenworth (Government Hill, 2; Engineer Hill, 1), 6; no locality more precise than county, 19. Saline County: Salina, Debold Farm, 4 (coll. of A. J. Kirn). Morris County: 1-1/2 mi. N Council Grove, 3. Douglas County: 1 mi. NW Midland, 2; 1 mi. N Lawrence, 1; 2-1/2 mi. W Lawrence, 2; 1 mi. W K. U. Campus, 2; 1 mi. W Lawrence, 2; 1/2 mi. W Lawrence, 2; "W K. U. Campus," 2; K. U. Campus, 4; Lawrence, 23; South Lawrence, 1; 1/2 mi. SW K. U. Campus, 2; Southwest K. U. Campus, 1; Haskell Institute, 1; 4-1/2 mi. S Lawrence, 1; 7 mi. SW Lawrence, 6; 7-1/2 mi. SW Lawrence, 1; 8 mi. SW Lawrence, 1; 10 mi. S Lawrence, 1; 11 mi. SW Lawrence, 3; no locality more precise than county, 15. Marion County: 1-1/2 mi. NE Lincolnville, 6; 4 mi. SE Lincolnville, 1; 6 mi. S Lincolnville, 1. Greenwood County: Hamilton, 1; 1/2 mi. S Hamilton, 4; 1 mi. S Hamilton, 4; 4 mi. S and 14 mi. W Hamilton, 6; 8 mi. SW Toronto, 1; 8-1/2 mi. SW Toronto, 5; no locality more precise than county, 6.
Geomys bursarius jugossicularis Hooper
Geomys lutescens jugossicularis Hooper, Occas. Papers Mus. Zoöl., Univ. Michigan, no. 420: 1, June 28, 1940; Hibbard, Trans. Kansas Acad. Sci., vol. 47, p. 75, 1944.
Type locality.—Lamar, Prowers County, Colorado.
Distribution in Kansas.—Extreme southwestern part of state, northward certainly to Hamilton County and south certainly to Morton and Seward counties.
Description.—A yellowish-cinnamon colored animal, with body of medium size, zygomatic plate of maxilla deep and mastoid process small.
Comparisons.—Differs from Geomys bursarius industrius in slightly lighter color; occiput not strongly inclined anterodorsally.