Inheritance of Characteristics in Domestic Fowl - Charles Benedict Davenport

Table 60.

Color.	In 64, calculated.	In the actual 85 individuals.
Color.	In 64, calculated.	Calculated.	Observed.
White.	52	69	68
Game.	9	12	16
Black.	3	4	1

The proportion of whites agrees closely with expectation. If this is not the case with the other two classes, the discrepancy must be attributed in part to insufficient observations and in part to the difficulties of precise classification in the early stages. The result is so close, however, as to lend strong support to our hypothesis as to the gametic constitution of the parents. Nothing is more striking, and to the unprejudiced mind more convincing, than the appearance of typically Game-colored birds in the grandchildren of wholly white parents.

3. SILKIE × BUFF COCHIN.
(Plates [7], [8].)

By hypothesis this cross is between cJnwx and CjnwX. The first generation should give the zygotic formula CcJjn₂w₂Xx, or, more simply, CcJjXx. The formula differs much from that of either parent, and the progeny themselves are no less remarkable. They have a washed-out buff color (since they are heterozygous in both C and X), and the Jungle pattern shows itself in the black tail and slightly redder buff of the wing-bar and hackles in the male. Since the hybrids are heterozygous in respect to 3 pairs of characters, when segregation occurs each parent produces 8 kinds of gametes, as follows: CJX, CJx, CjX, Cjx, cJX, cJx, cjX, cjx. In F₂ the types listed in table 61 may be expected in 64 offspring.

Table 61.—Distribution of colors, theoretic classes.—Probable frequency in 64 progeny.

Zygotic formula.	White.	Buff.	Buff + black.	Game.
C₂J₂X₂	...	...	1	...
C₂J₂Xx	...	...	2	...
C₂J₂x₂	...	...	...	1
C₂JjX₂	...	...	2	...
C₂JjXx	...	...	4	...
C₂Jjx₂	...	...	...	2
C₂j₂X₂	...	1	...	...
C₂j₂Xx	...	2	...	...
C₂j₂x₂	1	...	...	...
CcJ₂X₂	...	...	2	...
CcJ₂Xx	...	...	4	...
CcJ₂x₂	...	...	...	2
CcJjX₂	...	...	4	...
CcJjXx	...	...	8	...
CcJjx₂	...	...	...	4
Ccj₂x₂	...	2	...	...
Ccj₂Xx	...	4	...	...
Ccj₂x₂	2	...	...	...
c₂—	16	...	...	...
Total	19	9	27	9

The classification here employed can not be used in detail in comparing observed results with expectation, for the distinction between buff and buff-and-black appears only in chicks that have acquired the permanent plumage. Consequently it will be found necessary to combine these two classes into one and then make the comparison—as is done in table 62.

Table 62.—Distribution of colors, combined classes.

Color.	In 64, calculated.	In the actual 58 individuals.
Color.	In 64, calculated.	Calculated.	Observed.
Buff (and black).	36	33	34
White.	19	17	17
Game.	9	8	7
Total.	64	58	58

3. SILKIE × BUFF COCHIN.(Plates [7], [8].)

3. SILKIE × BUFF COCHIN.
(Plates [7], [8].)