Terga, irregularly oval, with the scutal margin straight; basal point blunt, with the two sides placed at about an angle of 45° to each other; the lower part of the carinal margin, immediately over the latera, (as seen internally,) is slightly hollowed out. Exteriorly, towards the bottom of the valve, from the overlapping of the scuta, of the latera, and of the carina, only a narrow rounded ridge is exposed, which runs down to the basal angle at about one third of the entire width of the valve, from the scutal margin. Internally the valve is slightly concave.
The Carina slightly overlaps the terga; internally concave; generally with a large upper portion freely projecting; inwardly curved, without any central crest or ridge; valve nearly as wide as the middle part of the terga; inner growing or corium-covered surface, with its basal margin, protuberant and arched.
Rostrum ([Pl. VIII], [fig. 1 a´, a], and greatly magnified [1 b´]) very narrow; rarely more than two or three layers of growth are preserved; the sides are deeply sinuous, owing to each zone widening downwards; basal margin rounded; in width equalling about two and a half of the uppermost scales of the peduncle, and about half as wide as the latera.
Latera, small, placed obliquely, and parallel to the lower carinal margin of the terga; longer axis equal to five of the uppermost scales of the peduncle, and to nearly half the width of the base of the carina; growing surface (or a section made parallel to the growth-layers,) is narrow, elliptic, pointed at both ends, but the carinal half rather thicker than the scutal half.
The Peduncle varies in length, generally about twice as long as the capitulum, in one specimen above thrice as long. The upper part as wide as the capitulum, the lower part sometimes much attenuated. The calcified scales in the uppermost whorl ([Pl. VIII], [fig. 1 b´]) are only slightly larger than those in the second whorl; the scales in the succeeding three or four whorls, are considerably larger than those below, which latter very gradually decrease in size, till, low down on the peduncle, they are barely visible to the naked eye. In this lower part, they may be called calcareous beads; they stand some way apart from each other; they are nearly hemispherical, smooth, translucent, and furnished with a conical fang; some of the smallest were 1/325th and 1/400th of an inch in diameter. The upper scales vary somewhat in the outline, the most usual shape being sub-triangular, with the lower margin arched and protuberant; and this margin, in the two or three upper whorls, is crenated with teeth, which are conical and sharp, after exuviation, but soon become reduced to mere notches. The scales in the uppermost whorl are usually nearly quadrilateral; the imbedded portion, or fang of each scale, is, in all, produced into a blunt rounded point. The basal calcareous cup ([fig. 1 a´] and [1 c´]) is well developed, and is sometimes even half an inch in diameter. Before the cup is formed, there is a row of small, flat discs ([fig. 1], and like those in [fig. 2 a´]) attached to the sides of the burrow: but a full account of these parts of the peduncle, and of the burrowing habits of this species, has been given under the generic description.
Size and Colour.—Full average-sized specimens have a capitulum half an inch in width and height; the entire length, with the contracted peduncle, being about an inch and a half. Valves coloured dirty white, with the enveloping membrane, when preserved, yellow. The outer maxillæ, palpi, pedicels of the cirri, anterior faces of the segments, dorsal tufts, caudal appendages, and penis, dark purple. Thoracic segments brown. There is a purple spot between the bases of the first pair of cirri.
Mouth.—Labrum considerably bullate, equalling about half the longitudinal diameter of the mouth; inferior part produced so as to separate the mouth some way from the adductor muscle; crest with a row of blunt teeth and hairs; central part depressed and flattened.
Palpi, rather large, separated from each other by only half their own length; bluntly pointed, thickly clothed with spines.
Mandibles ([Pl. X], [fig. 2]), with twice as many pectinations, namely 15, between the first and second main teeth, as between the second and third teeth, namely about 7; inferior angle strongly and coarsely pectinated; distance between the tips of the first and second main teeth, considerably less than between the tips of the second tooth and of the inferior angle; sides hirsute.
Maxillæ ([fig. 10]), with the edge not quite straight, with the whole inferior part slightly projecting; spines very numerous, thirty or forty pairs; those close beneath the two upper great unequal spines, form a tuft and are rather thinner than the others, as are also those near the inferior angle; sides hirsute.