Geological Observations on South America

by Charles Darwin

Professor Ehrenberg has examined for me specimens of the finer matter from in contact with these mammiferous remains: he finds in them two Polygastrica, decidedly marine forms; and six Phytolitharia, of which one is probably marine, and the others either of fresh-water or terrestrial origin. (“Monatsberichten der Akad. zu Berlin” April 1845. The list consists of:—

POLYGASTRICA. Gallionella sulcata. Stauroptera aspera? fragm.

PHYTOLITHARIA. Lithasteriscus tuberculatus. Lithostylidium Clepsammidium. Lithostylidium quadratum. Lithostylidium rude. Lithostylidium unidentatum. Spongolithis acicularis.)

Only one of these eight microscopical bodies is common to the nine from Monte Hermoso: but five of them are in common with those from the Pampean mud on the banks of the Parana. The presence of any fresh-water infusoria, considering the aridity of the surrounding country, is here remarkable: the most probable explanation appears to be, that these microscopical organisms were washed out of the adjoining great Pampean formation during its denudation, and afterwards redeposited.

We will now see what conclusions may be drawn from the facts above detailed. It is certain that the gravel-beds and intermediate red mud were deposited within the period, when existing species of Mollusca held to each other nearly the same relative proportions as they do on the present coast. These beds, from the number of littoral species, must have been accumulated in shallow water; but not, judging from the stratification of the gravel and the layers of marl, on a beach. From the manner in which the red clay fills up furrows in the underlying gravel, and is in some parts itself furrowed by the overlying gravel, whilst in other parts it either insensibly passes into, or alternates with, this upper gravel, we may infer several local changes in the currents, perhaps caused by slight changes, up or down, in the level of the land. By the elevation of these beds, to which period the alluvial mantle with pumice-pebbles, land and sea-shells belongs, the plain of Punta Alta, from twenty to thirty feet in height, was formed. In this neighbourhood there are other and higher sea-formed plains and lines of cliffs in the Pampean formation worn by the denuding action of the waves at different levels. Hence we can easily understand the presence of rounded masses of tosca-rock in this lowest plain; and likewise, as the cliffs at Monte Hermoso with their mammiferous remains stand at a higher level, the presence of the one much-rolled fragment of bone which was as black as jet: possibly some few of the other much-rolled bones may have been similarly derived, though I saw only the one fragment, in the same condition with those from Monte Hermoso. M. d’Orbigny has suggested that all these mammiferous remains may have been washed out of the Pampean formation, and afterwards redeposited together with the recent shells. (“Voyage” Part. Geolog. page 49.) Undoubtedly it is a marvellous fact that these numerous gigantic quadrupeds, belonging, with the exception of the Equus curvidens, to seven extinct genera, and one, namely, the Toxodon, not falling into any existing family, should have co-existed with Mollusca, all of which are still living species; but analogous facts have been observed in North America and in Europe. In the first place, it should not be overlooked, that most of the co-embedded shells have a more ancient and altered appearance than the bones. In the second place, is it probable that numerous bones not hardened by silex or any other mineral, could have retained their delicate prominences and surfaces perfect if they had been washed out of one deposit, and re-embedded in another:—this later deposit being formed of large, hard pebbles, arranged by the action of currents or breakers in shallow water into variously curved and inclined layers? The bones which are now in so perfect a state of preservation, must, I conceive, have been fresh and sound when embedded, and probably were protected by skin, flesh, or ligaments. The skeleton of the Scelidotherium indisputably was deposited entire: shall we say that when held together by its matrix it was washed out of an old gravel-bed (totally unlike in character to the Pampean formation), and re-embedded in another gravel-bed, composed (I speak after careful comparison) of exactly the same kind of pebbles, in the same kind of cement? I will lay no stress on the two cases of several ribs and bones of the extremities having APPARENTLY been embedded in their proper relative position: but will any one be so bold as to affirm that it is possible, that a piece of the thin tessellated armour of a Dasypoid quadruped, at least three feet long and two in width, and now so tender that I was unable with the utmost care to extract a fragment more than two or three inches square, could have been washed out of one bed, and re-embedded in another, together with some of the small bones of the feet, without having been dashed into atoms? We must then wholly reject M. d’Orbigny’s supposition, and admit as certain, that the Scelidotherium and the large Dasypoid quadruped, and as highly probable, that the Toxodon, Megatherium, etc., some of the bones of which are perfectly preserved, were embedded for the first time, and in a fresh condition, in the strata in which they were found entombed. These gigantic quadrupeds, therefore, though belonging to extinct genera and families, coexisted with the twenty above-enumerated Mollusca, the barnacle and two corals, still living on this coast. From the rolled fragment of black bone, and from the plain of Punta Alta being lower than that of Monte Hermoso, I conclude that the coarse sub-littoral deposits of Punta Alta, are of subsequent origin to the Pampean mud of Monte Hermoso; and the beds at this latter place, as we have seen, are probably of subsequent origin to the high tosca-plain round the Sierra Ventana: we shall, however, return, at the end of this chapter, to the consideration of these several stages in the great Pampean formation.

BUENOS AYRES TO ST. FE BAJADA, IN ENTRE RIOS.

For some distance northward of Buenos Ayres, the escarpment of the Pampean formation does not approach very near to the Plata, and it is concealed by vegetation: but in sections on the banks of the Rios Luxan, Areco, and Arrecifes, I observed both pale and dark reddish Pampean mud, with small, whitish concretions of tosca; at all these places mammiferous remains have been found. In the cliffs on the Parana, at San Nicolas, the Pampean mud contains but little tosca; here M. d’Orbigny found the remains of two rodents (Ctenomys Bonariensis and Kerodon antiquus) and the jaw of a Canis: when on the river I could clearly distinguish in this fine line of cliffs, “horizontal lines of variation both in tint and compactness.” (I quote these words from my note-book, as written down on the spot, on account of the general absence of stratification in the Pampean formation having been insisted on by M. d’Orbigny as a proof of the diluvial origin of this great deposit.) The plain northward of this point is very level, but with some depressions and lakes; I estimated its height at from forty to sixty feet above the Parana. At the A. Medio the bright red Pampean mud contains scarcely any tosca-rock; whilst at a short distance the stream of the Pabon, forms a cascade, about twenty feet in height, over a cavernous mass of two varieties of tosca-rock; of which one is very compact and semi- crystalline, with seams of crystallised carbonate of lime: similar compact varieties are met with on the Salidillo and Seco. The absolute identity (I speak after a comparison of my specimens) between some of these varieties, and those from Tapalguen, and from the ridge south of Bahia Blanca, a distance of 400 miles of latitude, is very striking.

At Rosario there is but little tosca-rock: near this place I first noticed at the edge of the river traces of an underlying formation, which, twenty- five miles higher up in the estancia of Gorodona, consists of a pale yellowish clay, abounding with concretionary cylinders of a ferruginous sandstone. This bed, which is probably the equivalent of the older tertiary marine strata, immediately to be described in Entre Rios, only just rises above the level of the Parana when low. The rest of the cliff at Gorodona, is formed of red Pampean mud, with, in the lower part, many concretions of tosca, some stalacti-formed, and with only a few in the upper part: at the height of six feet above the river, two gigantic skeletons of the Mastodon Andium were here embedded; their bones were scattered a few feet apart, but many of them still held their proper relative positions: they were much decayed and as soft as cheese, so that even one of the great molar teeth fell into pieces in my hand. We here see that the Pampean deposit contains mammiferous remains close to its base. On the banks of the Carcarana, a few miles distant, the lowest bed visible was pale Pampean mud, with masses of tosca-rock, in one of which I found a much decayed tooth of the Mastodon: above this bed, there was a thin layer almost composed of small concretions of white tosca, out of which I extracted a well preserved, but slightly broken tooth of Toxodon Platensis: above this there was an unusual bed of very soft impure sandstone. In this neighbourhood I noticed many single embedded bones, and I heard of others having been found in so perfect a state that they were long used as gate-posts: the Jesuit Falkner found here the dermal armour of some gigantic Edental quadruped.

In some of the red mud scraped from a tooth of one of the Mastodons at Gorodona, Professor Ehrenberg finds seven Polygastrica and thirteen Phytolitharia, all of them, I believe, with two exceptions, already known species. (“Monatsberichten der konig. Akad. zu Berlin” April 1845. The list consists of:—

POLYGASTRICA. Campylodiscus clypeus. Coscinodiscus subtilis. Coscinodiscus al. sp. Eunotia. Gallionella granulata. Himantidium gracile. Pinnularia borealis.)