A second theory, first suggested, I believe, by Sir W. Parish, is that the Pampean formation was thrown down on low and marshy plains by the rivers of this country before they assumed their present courses. The appearance and composition of the deposit, the manner in which it slopes up and round the primary ranges, the nature of the underlying marine beds, the estuary and sea-shells on the surface, the overlying sandstone beds at M. Hermoso, are all quite opposed to this view. Nor do I believe that there is a single instance of a skeleton of one of the extinct mammifers having been found in an upright position, as if it had been mired.

The third theory, of the truth of which I cannot entertain the smallest doubt, is that the Pampean formation was slowly accumulated at the mouth of the former estuary of the Plata and in the sea adjoining it. I have come to this conclusion from the reasons assigned against the two foregoing theories, and from simple geographical considerations. From the numerous shells of the Azara labiata lying loose on the surface of the plains, and near Buenos Ayres embedded in the tosca-rock, we know that this formation not only was formerly covered by, but that the uppermost parts were deposited in, the brackish water of the ancient La Plata. Southward and seaward of Buenos Ayres, the plains were upheaved from under water inhabited by true marine shells. We further know from Professor Ehrenberg’s examination of the twenty microscopical organisms in the mud round the tooth of the Mastodon high up the course of the Parana, that the bottom- most part of this formation was of brackish-water origin. A similar conclusion must be extended to the beds of like composition, at the level of the sea and under it, at M. Hermoso in Bahia Blanca. Dr. Carpenter finds that the harder varieties of tosca-rock, collected chiefly to the south, contain marine spongoid bodies, minute fragments of shells, corals, and Polythalamia; these perhaps may have been drifted inwards by the tides, from the more open parts of the sea. The absence of shells, throughout this deposit, with the exception of the uppermost layers near Buenos Ayres, is a remarkable fact: can it be explained by the brackish condition of the water, or by the deep mud at the bottom? I have stated that both the reddish mud and the concretions of tosca-rock are often penetrated by minute, linear cavities, such as frequently may be observed in fresh-water calcareous deposits:—were they produced by the burrowing of small worms? Only on this view of the Pampean formation having been of estuary origin, can the extraordinary numbers (presently to be alluded to) of the embedded mammiferous remains be explained. (It is almost superfluous to give the numerous cases (for instance, in Sumatra; Lyell “Principles” volume 3 page 325 sixth edition, of the carcasses of animals having been washed out to sea by swollen rivers; but I may refer to a recent account by Mr. Bettington “Asiatic Society” 1845 June 21st, of oxen, deer, and bears being carried into the Gulf of Cambray; see also the account in my “Journal” 2nd edition page 133, of the numbers of animals drowned in the Plata during the great, often recurrent, droughts.)

With respect to the first origin of the reddish mud, I will only remark, that the enormous area of Brazil consists in chief part of gneissic and other granitic rocks, which have suffered decomposition, and been converted into a red, gritty, argillaceous mass, to a greater depth than in any other country which I have seen. The mixture of rounded grains, and even of small fragments and pebbles of quartz, in the Pampean mud of Banda Oriental, is evidently due to the neighbouring and underlying primary rocks. The estuary mud was drifted during the Pampean period in a much more southerly course, owing probably to the east and west primary ridges south of the Plata not having been then elevated, than the mud of the Plata at present is; for it was formerly deposited as far south as the Colorado. The quantity of calcareous matter in this formation, especially in those large districts where the whole mass passes into tosca-rock, is very great: I have already remarked on the close resemblance in external and microscopical appearance, between this tosca-rock and the strata at Coquimbo, which have certainly resulted from the decay and attrition of recent shells: I dare not, however, extend this conclusion to the calcareous rocks of the Pampas, more especially as the underlying tertiary strata in western Banda Oriental show that at that period there was a copious emission of carbonate of lime, in connection with volcanic action. (I may add, that there are nearly similar superficial calcareous beds at King George’s Sound in Australia; and these undoubtedly have been formed by the disintegration of marine remains see “Volcanic Islands” etc. page 144. There is, however, something very remarkable in the frequency of superficial, thin beds of earthy calcareous matter, in districts where the surrounding rocks are not calcareous. Major Charters, in a Paper read before the Geographical Society April 13, 1840 and abstracted in the “Athenaeum” page 317, states that this is the case in parts of Mexico, and that he has observed similar appearances in many parts of South Africa. The circumstance of the uppermost stratum round the ragged Sierra Ventana, consisting of calcareous or marly matter, without any covering of alluvial matter, strikes me as very singular, in whatever manner we view the deposition and elevation of the Pampean formation.)

The Pampean formation, judging from its similar composition, and from the apparent absolute specific identity of some of its mammiferous remains, and from the generic resemblance of others, belongs over its vast area— throughout Banda Oriental, Entre Rios, and the wide extent of the Pampas as far south as the Colorado,—to the same geological epoch. The mammiferous remains occur at all depths from the top to the bottom of the deposit; and I may add that nowhere in the Pampas is there any appearance of much superficial denudation: some bones which I found near the Guardia del Monte were embedded close to the surface; and this appears to have been the case with many of those discovered in Banda Oriental: on the Matanzas, twenty miles south of Buenos Ayres, a Glyptodon was embedded five feet beneath the surface; numerous remains were found by S. Muniz, near Luxan, at an average depth of eighteen feet; in Buenos Ayres a skeleton was disinterred at sixty feet depth, and on the Parana I have described two skeletons of the Mastodon only five or six feet above the very base of the deposit. With respect to the age of this formation, as judged of by the ordinary standard of the existence of Mollusca, the only evidence within the limits of the true Pampas which is at all trustworthy, is afforded by the still living Azara labiata being embedded in tosca-rock near Buenos Ayres. At Punta Alta, however, we have seen that several of the extinct mammifers, most characteristic of the Pampean formation, co-existed with twenty species of Mollusca, a barnacle and two corals, all still living on this same coast;— for when we remember that the shells have a more ancient appearance than the bones; that many of the bones, though embedded in a coarse conglomerate, are perfectly preserved; that almost all the parts of the skeleton of the Scelidotherium, even to the knee-cap, were lying in their proper relative positions; and that a large piece of the fragile dermal armour of a Dasypoid quadruped, connected with some of the bones of the foot, had been entombed in a condition allowing the two sides to be doubled together, it must assuredly be admitted that these mammiferous remains were embedded in a fresh state, and therefore that the living animals co-existed with the co-embedded shells. Moreover, the Macrauchenia Patachonica (of which, according to Professor Owen, remains also occur in the Pampas of Buenos Ayres, and at Punta Alta) has been shown by satisfactory evidence of another kind, to have lived on the plains of Patagonia long after the period when the adjoining sea was first tenanted by its present commonest molluscous animals. We must, therefore, conclude that the Pampean formation belongs, in the ordinary geological sense of the word, to the Recent Period. (M. d’Orbigny believes “Voyage” Part. Geolog. page 81, that this formation, though “tres voisine de la notre, est neanmoins de beaucoup anterieure a notre creation.”)

At St. Fe Bajada, the Pampean estuary formation, with its mammiferous remains, conformably overlies the marine tertiary strata, which (as first shown by M. d’Orbigny) are contemporaneous with those of Patagonia, and which, as we shall hereafter see, belong to a very ancient tertiary stage. When examining the junction between these two formations, I thought that the concretionary layer of marl marked a passage between the marine and estuary stages. M. d’Orbigny disputes this view (as given in my “Journal”), and I admit that it is erroneous, though in some degree excusable, from their conformability and from both abounding with calcareous matter. It would, indeed, have been a great anomaly if there had been a true passage between a deposit contemporaneous with existing species of mollusca, and one in which all the mollusca appear to be extinct. Northward of Santa Fe, M. d’Orbigny met with ferruginous sandstones, marly rocks, and other beds, which he considers as a distinct and lower formation; but the evidence that they are not parts of the same with an altered mineralogical character, does not appear to me quite satisfactory.

In Western Banda Oriental, while the marine tertiary strata were accumulating, there were volcanic eruptions, much silex and lime were precipitated from solution, coarse conglomerates were formed, being derived probably from adjoining land, and layers of red mud and marly rocks, like those of the Pampean formation, were occasionally deposited. The true Pampean deposit, with mammiferous remains, instead of as at Santa Fe overlying conformably the tertiary strata, is here seen at a lower level folding round and between the flat-topped, cliff-bounded hills, formed by a upheaval and denudation of these same tertiary strata. The upheaval, having occurred here earlier than at Santa Fe, may be naturally accounted for by the contemporaneous volcanic action. At the Barrancas de S. Gregorio, the Pampean deposit, as we have seen, overlies and fills up furrows in coarse sand, precisely like that now accumulating on the shores near the mouth of the Plata. I can hardly believe that this loose and coarse sand is contemporaneous with the old tertiary and often crystalline strata of the more western parts of the province; and am induced to suspect that it is of subsequent origin. If that section near Colonia could be implicitly trusted, in which, at a height of only fifteen feet above the Plata, a bed of fresh-looking mussels, of an existing littoral species, appeared to lie between the sand and the Pampean mud, I should conclude that Banda Oriental must have stood, when the coarse sand was accumulating, at only a little below its present level, and had then subsided, allowing the estuary Pampean mud to cover far and wide its surface up to a height of some hundred feet; and that after this subsidence the province had been uplifted to its present level.

In Western Banda Oriental, we know, from two unequivocal sections that there is a mass, absolutely undistinguishable from the true Pampean deposit, beneath the old tertiary strata. This inferior mass must be very much more ancient than the upper deposit with its mammiferous remains, for it lies beneath the tertiary strata in which all the shells are extinct. Nevertheless, the lower and upper masses, as well as some intermediate layers, are so similar in mineralogical character, that I cannot doubt that they are all of estuary origin, and have been derived from the same great source. At first it appeared to me extremely improbable, that mud of the same nature should have been deposited on nearly the same spot, during an immense lapse of time, namely, from a period equivalent perhaps to the Eocene of Europe to that of the Pampean formation. But as, at the very commencement of the Pampean period, if not at a still earlier period, the Sierra Ventana formed a boundary to the south,—the Cordillera or the plains in front of them to the west,—the whole province of Corrientes probably to the north, for, according to M. d’Orbigny, it is not covered by the Pampean deposit,—and Brazil, as known by the remains in the caves, to the north-east; and as again, during the older tertiary period, land already existed in Western Banda Oriental and near St. Fe Bajada, as may be inferred from the vegetable debris, from the quantities of silicified wood, and from the remains of a Toxodon found, according to M. d’Orbigny, in still lower strata, we may conclude, that at this ancient period a great expanse of water was surrounded by the same rocky framework which now bounds the plains of Pampean formation. This having been the case, the circumstance of sediment of the same nature having been deposited in the same area during an immense lapse of time, though highly remarkable, does not appear incredible.

The elevation of the Pampas, at least of the southern parts, has been slow and interrupted by several periods of rest, as may be inferred from the plains, cliffs, and lines of sand-dunes (with shells and pumice-pebbles) standing at different heights. I believe, also, that the Pampean mud continued to be deposited, after parts of this formation had already been elevated, in the same manner as mud would continue to be deposited in the estuary of the Plata, if the mud-banks on its shores were now uplifted and changed into plains: I believe in this from the improbability of so many skeletons and bones having been accumulated at one spot, where M. Hermoso now stands, at a depth of between eight hundred and one thousand feet, and at a vast distance from any land except small rocky islets,—as must have been the case, if the high tosca-plain round the Ventana and adjoining Sierras, had not been already uplifted and converted into land, supporting mammiferous animals. At Punta Alta we have good evidence that the gravel- strata, which certainly belong to the true Pampean period, were accumulated after the elevation in that neighbourhood of the main part of the Pampean deposit, whence the rounded masses of tosca-rock were derived, and that rolled fragment of black bone in the same peculiar condition with the remains at Monte Hermoso.

The number of the mammiferous remains embedded in the Pampas is, as I have remarked, wonderful: it should be borne in mind that they have almost exclusively been found in the cliffs and steep banks of rivers, and that, until lately, they excited no attention amongst the inhabitants: I am firmly convinced that a deep trench could not be cut in any line across the Pampas, without intersecting the remains of some quadruped. It is difficult to form an opinion in what part of the Pampas they are most numerous; in a limited spot they could not well have been more numerous than they were at P. Alta; the number, however, lately found by Senor F. Muniz, near Luxan, in a central spot in the Pampas, is extraordinarily great: at the end of this chapter I will give a list of all the localities at which I have heard of remains having been discovered. Very frequently the remains consist of almost perfect skeletons; but there are, also, numerous single bones, as for instance at St. Fe. Their state of preservation varies much, even when embedded near each other: I saw none others so perfectly preserved as the heads of the Toxodon and Mylodon from the white soft earthy bed on the Sarandis in Banda Oriental. It is remarkable that in two limited sections I found no less than five teeth separately embedded, and I heard of teeth having been similarly found in other parts: may we suppose that the skeletons or heads were for a long time gently drifted by currents over the soft muddy bottom, and that the teeth occasionally, here and there, dropped out?

It may be naturally asked, where did these numerous animals live? From the remarkable discoveries of MM. Lund and Clausen, it appears that some of the species found in the Pampas inhabited the highlands of Brazil: the Mastodon Andium is embedded at great heights in the Cordillera from north of the equator to at least as far south as Tarija (Humboldt states that the Mastodon has been discovered in New Granada: it has been found in Quito. When at Lima, I saw a tooth of a Mastodon in the possession of Don M. Rivero, found at Playa Chica on the Maranon, near the Guallaga. Every one has heard of the numerous remains of Mastodon in Bolivia.); and as there is no higher land, there can be little doubt that this Mastodon must have lived on the plains and valleys of that great range. These countries, however, appear too far distant for the habitation of the individuals entombed in the Pampas: we must probably look to nearer points, for instance to the province of Corrientes, which, as already remarked, is said not to be covered by the Pampean formation, and may therefore, at the period of its deposition, have existed as dry land. I have already given my reasons for believing that the animals embedded at M. Hermoso and at P. Alta in Bahia Blanca, lived on adjoining land, formed of parts of the already elevated Pampean deposit. With respect to the food of these many great extinct quadrupeds, I will not repeat the facts given in my “Journal” (second edition page 85), showing that there is no correlation between the luxuriance of the vegetation of a country and the size of its mammiferous inhabitants. I do not doubt that large animals could now exist, as far as the amount, not kind, of vegetation is concerned, on the sterile plains of Bahia Blanca and of the R. Negro, as well as on the equally, if not more sterile plains of Southern Africa. The climate, however, may perhaps have somewhat deteriorated since the mammifers embedded at Bahia Blanca lived there; for we must not infer, from the continued existence of the same shells on the present coasts, that there has been no change in climate; for several of these shells now range northward along the shores of Brazil, where the most luxuriant vegetation flourishes under a tropical temperature. With respect to the extinction, which at first fills the mind with astonishment, of the many great and small mammifers of this period, I may also refer to the work above cited (second edition page 173), in which I have endeavoured to show, that however unable we may be to explain the precise cause, we ought not properly to feel more surprised at a species becoming extinct than at one being rare; and yet we are accustomed to view the rarity of any particular species as an ordinary event, not requiring any extraordinary agency.