Dear Professor Semper,
Since writing I have recalled some of the thoughts and conclusions which have passed through my mind of late years. In North America, in going from north to south or from east to west, it is clear that the changed conditions of life have modified the organisms in the different regions, so that they now form distinct races or even species. It is further clear that in isolated districts, however small, the inhabitants almost always get slightly modified, and how far this is due to the nature of the slightly different conditions to which they are exposed, and how far to mere interbreeding, in the manner explained by Weismann, I can form no opinion. The same difficulty occurred to me (as shown in my 'Variation of Animals and Plants under Domestication') with respect to the aboriginal breeds of cattle, sheep, etc., in the separated districts of Great Britain, and indeed throughout Europe. As our knowledge advances, very slight differences, considered by systematists as of no importance in structure, are continually found to be functionally important; and I have been especially struck with this fact in the case of plants to which my observations have of late years been confined. Therefore it seems to me rather rash to consider the slight differences between representative species, for instance those inhabiting the different islands of the same archipelago, as of no functional importance, and as not in any way due to natural selection. With respect to all adapted structures, and these are innumerable, I cannot see how M. Wagner's view throws any light, nor indeed do I see at all more clearly than I did before, from the numerous cases which he has brought forward, how and why it is that a long isolated form should almost always become slightly modified. I do not know whether you will care about hearing my further opinion on the point in question, for as before remarked I have not attended much of late years to such questions, thinking it prudent, now that I am growing old, to work at easier subjects.
Believe me, yours very sincerely, CH. DARWIN.
I hope and trust that you will throw light on these points.
P.S.—I will add another remark which I remember occurred to me when I first read M. Wagner. When a species first arrives on a small island, it will probably increase rapidly, and unless all the individuals change instantaneously (which is improbable in the highest degree), the slowly, more or less, modifying offspring must intercross one with another, and with their unmodified parents, and any offspring not as yet modified. The case will then be like that of domesticated animals which have slowly become modified, either by the action of the external conditions or by the process which I have called the UNCONSCIOUS SELECTION by man—i.e., in contrast with methodical selection.
[The letters continue the history of the year 1872, which has been interrupted by a digression on Isolation.]
CHARLES DARWIN TO THE MARQUIS DE SAPORTA. Down, April 8, 1872.
Dear Sir,
I thank you very sincerely and feel much honoured by the trouble which you have taken in giving me your reflections on the origin of Man. It gratifies me extremely that some parts of my work have interested you, and that we agree on the main conclusion of the derivation of man from some lower form.
I will reflect on what you have said, but I cannot at present give up my belief in the close relationship of Man to the higher Simiae. I do not put much trust in any single character, even that of dentition; but I put the greatest faith in resemblances in many parts of the whole organisation, for I cannot believe that such resemblances can be due to any cause except close blood relationship. That man is closely allied to the higher Simiae is shown by the classification of Linnaeus, who was so good a judge of affinity. The man who in England knows most about the structure of the Simiae, namely, Mr. Mivart, and who is bitterly opposed to my doctrines about the derivation of the mental powers, yet has publicly admitted that I have not put man too close to the higher Simiae, as far as bodily structure is concerned. I do not think the absence of reversions of structure in man is of much weight; C. Vogt, indeed, argues that [the existence of] Micr-cephalous idiots is a case of reversion. No one who believes in Evolution will doubt that the Phocae are descended from some terrestrial Carnivore. Yet no one would expect to meet with any such reversion in them. The lesser divergence of character in the races of man in comparison with the species of Simiadae may perhaps be accounted for by man having spread over the world at a much later period than did the Simiadae. I am fully prepared to admit the high antiquity of man; but then we have evidence, in the Dryopithecus, of the high antiquity of the Anthropomorphous Simiae.