I have expressly stated that I believe physical conditions have a more direct effect on plants than on animals. But the more I study, the more I am led to think that natural selection regulates, in a state of nature, most trifling differences. As squared stone, or bricks, or timber, are the indispensable materials for a building, and influence its character, so is variability not only indispensable, but influential. Yet in the same manner as the architect is the ALL important person in a building, so is selection with organic bodies...
[The meeting of the British Association at Oxford in 1860 is famous for two pitched battles over the 'Origin of Species.' Both of them originated in unimportant papers. On Thursday, June 28, Dr. Daubeny of Oxford made a communication to Section D: "On the final causes of the sexuality of plants, with particular reference to Mr. Darwin's work on the 'Origin of Species.'" Mr. Huxley was called on by the President, but tried (according to the "Athenaeum" report) to avoid a discussion, on the ground "that a general audience, in which sentiment would unduly interfere with intellect, was not the public before which such a discussion should be carried on." However, the subject was not allowed to drop. Sir R. Owen (I quote from the "Athenaeum", July 7, 1860), who "wished to approach this subject in the spirit of the philosopher," expressed his "conviction that there were facts by which the public could come to some conclusion with regard to the probabilities of the truth of Mr. Darwin's theory." He went on to say that the brain of the gorilla "presented more differences, as compared with the brain of man, than it did when compared with the brains of the very lowest and most problematical of the Quadrumana." Mr. Huxley replied, and gave these assertions a "direct and unqualified contradiction," pledging himself to "justify that unusual procedure elsewhere" ('Man's Place in Nature,' by T.H. Huxley, 1863, page 114.), a pledge which he amply fulfilled. (See the 'Nat. Hist. Review,' 1861.) On Friday there was peace, but on Saturday 30th, the battle arose with redoubled fury over a paper by Dr. Draper of New York, on the 'Intellectual development of Europe considered with reference to the views of Mr. Darwin.'
The following account is from an eye-witness of the scene.
"The excitement was tremendous. The Lecture-room, in which it had been arranged that the discussion should be held, proved far too small for the audience, and the meeting adjourned to the Library of the Museum, which was crammed to suffocation long before the champions entered the lists. The numbers were estimated at from 700 to 1000. Had it been term-time, or had the general public been admitted, it would have been impossible to have accommodated the rush to hear the oratory of the bold Bishop. Professor Henslow, the President of Section D, occupied the chair and wisely announced in limine that none who had not valid arguments to bring forward on one side or the other, would be allowed to address the meeting: a caution that proved necessary, for no fewer than four combatants had their utterances burked by him, because of their indulgence in vague declamation.
"The Bishop was up to time, and spoke for full half-an-hour with inimitable spirit, emptiness and unfairness. It was evident from his handling of the subject that he had been 'crammed' up to the throat, and that he knew nothing at first hand; in fact, he used no argument not to be found in his 'Quarterly' article. He ridiculed Darwin badly, and Huxley savagely, but all in such dulcet tones, so persuasive a manner, and in such well-turned periods, that I who had been inclined to blame the President for allowing a discussion that could serve no scientific purpose now forgave him from the bottom of my heart. Unfortunately the Bishop, hurried along on the current of his own eloquence, so far forgot himself as to push his attempted advantage to the verge of personality in a telling passage in which he turned round and addressed Huxley: I forgot the precise words, and quote from Lyell. 'The Bishop asked whether Huxley was related by his grandfather's or grandmother's side to an ape.' (Lyell's 'Letters,' vol. ii. page 335.) Huxley replied to the scientific argument of his opponent with force and eloquence, and to the personal allusion with a sel-restraint, that gave dignity to his crushing rejoinder."
Many versions of Mr. Huxley's speech were current: the following report of his conclusion is from a letter addressed by the late John Richard Green, then an undergraduate, to a fellow-student, now Professor Boyd Dawkins. "I asserted, and I repeat, that a man has no reason to be ashamed of having an ape for his grandfather. If there were an ancestor whom I should feel shame in recalling, it would be a MAN, a man of restless and versatile intellect, who, not content with an equivocal (Prof. V. Carus, who has a distinct recollection of the scene, does not remember the word equivocal. He believes too that Lyell's version of the "ape" sentence is slightly incorrect.) success in his own sphere of activity, plunges into scientific questions with which he has no real acquaintance, only to obscure them by an aimless rhetoric, and distract the attention of his hearers from the real point at issue by eloquent digressions, and skilled appeals to religious prejudice."
The letter above quoted continues:
"The excitement was now at its height; a lady fainted and had to be carried out, and it was some time before the discussion was resumed. Some voices called for Hooker, and his name having been handed up, the President invited him to give his view of the theory from the Botanical side. This he did, demonstrating that the Bishop, by his own showing, had never grasped the principles of the 'Origin' (With regard to the Bishop's 'Quarterly Review,' my father wrote: "These very clever men think they can write a review with a very slight knowledge of the book reviewed or subject in question."), and that he was absolutely ignorant of the elements of botanical science. The Bishop made no reply, and the meeting broke up.
"There was a crowded conversazione in the evening at the rooms of the hospitable and genial Professor of Botany, Dr. Daubeny, where the almost sole topic was the battle of the 'Origin,' and I was much struck with the fair and unprejudiced way in which the black coats and white cravats of Oxford discussed the question, and the frankness with which they offered their congratulations to the winners in the combat.]
CHARLES DARWIN TO J.D. HOOKER. Sudbrook Park, Monday night [July 2nd, 1860].