LETTER 726. TO J.D. HOOKER. Down, August 30th {1874}.
I am particularly obliged for your address. (726/1. Presidential address (Biological Section) at the Belfast meeting of the British Association, 1874.) It strikes me as quite excellent, and has interested me in the highest degree. Nor is this due to my having worked at the subject, for I feel sure that I should have been just as much struck, perhaps more so, if I had known nothing about it. You could not, in my opinion, have put the case better. There are several lights (besides the facts) in your essay new to me, and you have greatly honoured me. I heartily congratulate you on so splendid a piece of work. There is a misprint at page 7, Mitschke for Nitschke. There is a partial error at page 8, where you say that Drosera is nearly indifferent to organic substances. This is much too strong, though they do act less efficiently than organic with soluble nitrogenous matter; but the chief difference is in the widely different period of subsequent re-expansion. Thirdly, I did not suggest to Sanderson his electrical experiments, though, no doubt, my remarks led to his thinking of them.
Now for your letter: you are very generous about Dionoea, but some of my experiments will require cutting off leaves, and therefore injuring plants. I could not write to Lady Dorothy {Nevill}. Rollisson says that they expect soon a lot from America. If Dionoea is not despatched, have marked on address, "to be forwarded by foot-messenger."
Mrs. Barber's paper is very curious, and ought to be published (726/2. Mrs. Barber's paper on the pupa of Papilio Nireus assuming different tints corresponding to the objects to which it was attached, was communicated by Mr. Darwin to the "Trans. Entomolog. Soc." 1874.); but when you come here (and REMEMBER YOU OFFERED TO COME) we will consult where to send it. Let me hear when you recommence on Cephalotus or Sarracenia, as I think I am now on right track about Utricularia, after wasting several weeks in fruitless trials and observations. The negative work takes five times more time than the positive.
LETTER 727. TO J.D. HOOKER. Down, September 18th {1874}.
I have had a splendid day's work, and must tell you about it.
Lady Dorothy sent me a young plant of U{tricularia} montana (727/1. See "Life and Letters," III., page 327, and "Insectivorous Plants," page 431.), which I fancy is the species you told me of. The roots or rhizomes (for I know not which they are; I can see no scales or internodes or absorbent hairs) bear scores of bladders from 1/20 to 1/100 of an inch in diameter; and I traced these roots to the depth of 1 1/2 in. in the peat and sand. The bladders are like glass, and have the same essential structure as those of our species, with the exception that many exterior parts are aborted. Internally the structure is perfect, as is the minute valvular opening into the bladder, which is filled with water. I then felt sure that they captured subterranean insects, and after a time I found two with decayed remnants, with clear proof that something had been absorbed, which had generated protoplasm. When you are here I shall be very curious to know whether they are roots or rhizomes.
Besides the bladders there are great tuber-like swellings on the rhizomes; one was an inch in length and half in breadth. I suppose these must have been described. I strongly suspect that they serve as reservoirs for water. (727/2. The existence of water-stores is quite in accordance with the epiphytic habit of the plant.) But I shall experimentise on this head. A thin slice is a beautiful object, and looks like coarsely reticulated glass.
If you have an old plant which could be turned out of its pot (and can spare the time), it would be a great gain to me if you would tear off a bit of the roots near the bottom, and shake them well in water, and see whether they bear these minute glass-like bladders. I should also much like to know whether old plants bear the solid bladder-like bodies near the upper surface of the pot. These bodies are evidently enlargements of the roots or rhizomes. You must forgive this long letter, and make allowance for my delight at finding this new sub-group of insect-catchers. Sir E. Tennent speaks of an aquatic species of Utricularia in Ceylon, which has bladders on its roots, and rises annually to the surface, as he says, by this means. (727/3. Utricularia stellaris. Emerson Tennent's "Ceylon," Volume I., page 124, 1859.)
We shall be delighted to see you here on the 26th; if you will let us know your train we will send to meet you. You will have to work like a slave while you are here.