We have seen in the first chapter that the homological structure of man, his embryological development and the rudiments which he still retains, all declare in the plainest manner that he is descended from some lower form. The possession of exalted mental powers is no insuperable objection to this conclusion. In order that an ape-like creature should have been transformed into man, it is necessary that this early form, as well as many successive links, should all have varied in mind and body. It is impossible to obtain direct evidence on this head; but if it can be shewn that man now varies—that his variations are induced by the same general causes, and obey the same general laws, as in the case of the lower animals—there can be little doubt that the preceding intermediate links varied in a like manner. The variations at each successive stage of descent must, also, have been in some manner accumulated and fixed.
The facts and conclusions to be given in this chapter relate almost exclusively to the probable means by which the transformation of man has been effected, as far as his bodily structure is concerned. The following chapter will be devoted to the development of his intellectual and moral faculties. But the present discussion likewise bears on the origin of the different races or species of mankind, whichever term may be preferred.
It is manifest that man is now subject to much variability. No two individuals of the same race are quite alike. We may compare millions of faces, and each will be distinct. There is an equally great amount of diversity in the proportions and dimensions of the various parts of the body; the length of the legs being one of the most variable points.[140] Although in some quarters of the world an elongated skull, and in other quarters a short skull prevails, yet there is great diversity of shape even within the limits of the same race, as with the aborigines of America and South Australia,—the latter a race “probably as pure and homogeneous in blood, customs, and language as any in existence”—and even with the inhabitants of so confined an area as the Sandwich Islands.[141] An eminent dentist assures me that there is nearly as much diversity in the teeth, as in the features. The chief arteries so frequently run in abnormal courses, that it has been found useful for surgical purposes to calculate from 12,000 corpses how often each course prevails.[142] The muscles are eminently variable: thus those of the foot were found by Prof. Turner[143] not to be strictly alike in any two out of fifty bodies; and in some the deviations were considerable. Prof. Turner adds that the power of performing the appropriate movements must have been modified in accordance with the several deviations. Mr. J. Wood has recorded[144] the occurrence of 295 muscular variations in thirty-six subjects, and in another set of the same number no less than 558 variations, reckoning both sides of the body as one. In the last set, not one body out of the thirty-six was “found totally wanting in departures from the standard descriptions of the muscular system given in anatomical text-books.” A single body presented the extraordinary number of twenty-five distinct abnormalities. The same muscle sometimes varies in many ways: thus Prof. Macalister describes[145] no less than twenty distinct variations in the palmaris accessorius.
The famous old anatomist, Wolff,[146] insists that the internal viscera are more variable than the external parts: Nulla particula est quæ non aliter et aliter in aliis se habeat hominibus. He has even written a treatise on the choice of typical examples of the viscera for representation. A discussion on the beau-ideal of the liver, lungs, kidneys, &c., as of the human face divine, sounds strange in our ears.
The variability or diversity of the mental faculties in men of the same race, not to mention the greater differences between the men of distinct races, is so notorious that not a word need here be said. So it is with the lower animals, as has been illustrated by a few examples in the last chapter. All who have had charge of menageries admit this fact, and we see it plainly in our dogs and other domestic animals. Brehm especially insists that each individual monkey of those which he kept under confinement in Africa had its own peculiar disposition and temper: he mentions one baboon remarkable for its high intelligence; and the keepers in the Zoological Gardens pointed out to me a monkey, belonging to the New World division, equally remarkable for intelligence. Rengger, also, insists on the diversity in the various mental characters of the monkeys of the same species which he kept in Paraguay; and this diversity, as he adds, is partly innate, and partly the result of the manner in which they have been treated or educated.[147]
I have elsewhere[148] so fully discussed the subject of Inheritance that I need here add hardly anything. A greater number of facts have been collected with respect to the transmission of the most trifling, as well as of the most important characters in man than in any of the lower animals; though the facts are copious enough with respect to the latter. So in regard to mental qualities, their transmission is manifest in our dogs, horses, and other domestic animals. Besides special tastes and habits, general intelligence, courage, bad and good temper, &c., are certainly transmitted. With man we see similar facts in almost every family; and we now know through the admirable labours of Mr. Galton[149] that genius, which implies a wonderfully complex combination of high faculties, tends to be inherited; and, on the other hand, it is too certain that insanity and deteriorated mental powers likewise run in the same families.
With respect to the causes of variability we are in all cases very ignorant; but we can see that in man as in the lower animals, they stand in some relation with the conditions to which each species has been exposed during several generations. Domesticated animals vary more than those in a state of nature; and this is apparently due to the diversified and changing nature of their conditions. The different races of man resemble in this respect domesticated animals, and so do the individuals of the same race when inhabiting a very wide area, like that of America. We see the influence of diversified conditions in the more civilised nations, the members of which belong to different grades of rank and follow different occupations, presenting a greater range of character than the members of barbarous nations. But the uniformity of savages has often been exaggerated, and in some cases can hardly be said to exist.[150] It is nevertheless an error to speak of man, even if we look only to the conditions to which he has been subjected, as “far more domesticated”[151] than any other animal. Some savage races, such as the Australians, are not exposed to more diversified conditions than are many species which have very wide ranges. In another and much more important respect, man differs widely from any strictly domesticated animal; for his breeding has not been controlled, either through methodical or unconscious selection. No race or body of men has been so completely subjugated by other men, that certain individuals have been preserved and thus unconsciously selected, from being in some way more useful to their masters. Nor have certain male and female individuals been intentionally picked out and matched, except in the well-known case of the Prussian grenadiers; and in this case man obeyed, as might have been expected, the law of methodical selection; for it is asserted that many tall men were reared in the villages inhabited by the grenadiers with their tall wives.
If we consider all the races of man, as forming a single species, his range is enormous; but some separate races, as the Americans and Polynesians, have very wide ranges. It is a well-known law that widely-ranging species are much more variable than species with restricted ranges; and the variability of man may with more truth be compared with that of widely-ranging species, than with that of domesticated animals.
Not only does variability appear to be induced in man and the lower animals by the same general causes, but in both the same characters are affected in a closely analogous manner. This has been proved in such full detail by Godron and Quatrefages, that I need here only refer to their works.[152] Monstrosities, which graduate into slight variations, are likewise so similar in man and the lower animals, that the same classification and the same terms can be used for both, as may be seen in Isidore Geoffroy St.-Hilaire’s great work.[153] This is a necessary consequence of the same laws of change prevailing throughout the animal kingdom. In my work on the variation of domestic animals, I have attempted to arrange in a rude fashion the laws of variation under the following heads:—The direct and definite action of changed conditions, as shewn by all or nearly all the individuals of the same species varying in the same manner under the same circumstances. The effects of the long-continued use or disuse of parts. The cohesion of homologous parts. The variability of multiple parts. Compensation of growth; but of this law I have found no good instances in the case of man. The effects of the mechanical pressure of one part on another; as of the pelvis on the cranium of the infant in the womb. Arrests of development, leading to the diminution or suppression of parts. The reappearance of long-lost characters through reversion. And lastly, correlated variation. All these so-called laws apply equally to man and the lower animals; and most of them even to plants. It would be superfluous here to discuss all of them;[154] but several are so important for us, that they must be treated at considerable length.
The direct and definite action of changed conditions.—This is a most perplexing subject. It cannot be denied that changed conditions produce some effect, and occasionally a considerable effect, on organisms of all kinds; and it seems at first probable that if sufficient time were allowed this would be the invariable result. But I have failed to obtain clear evidence in favour of this conclusion; and valid reasons may be urged on the other side, at least as far as the innumerable structures are concerned, which are adapted for special ends. There can, however, be no doubt that changed conditions induce an almost indefinite amount of fluctuating variability, by which the whole organisation is rendered in some degree plastic.