The anthropomorphous apes, namely the gorilla, chimpanzee, orang, and hylobates, are separated as a distinct sub-group from the other Old World monkeys by most naturalists. I am aware that Gratiolet, relying on the structure of the brain, does not admit the existence of this sub-group, and no doubt it is a broken one; thus the orang, as Mr. St. G. Mivart remarks,[266] “is one of the most peculiar and aberrant forms to be found in the Order.” The remaining, non-anthropomorphous, Old World monkeys, are again divided by some naturalists into two or three smaller sub-groups; the genus Semnopithecus, with its peculiar sacculated stomach, being the type of one such sub-group. But it appears from M. Gaudry’s wonderful discoveries in Attica, that during the Miocene period a form existed there, which connected Semnopithecus and Macacus; and this probably illustrates the manner in which the other and higher groups were once blended together.
If the anthropomorphous apes be admitted to form a natural sub-group, then as man agrees with them, not only in all those characters which he possesses in common with the whole Catarhine group, but in other peculiar characters, such as the absence of a tail and of callosities and in general appearance, we may infer that some ancient member of the anthropomorphous sub-group gave birth to man. It is not probable that a member of one of the other lower sub-groups should, through the law of analogous variation, have given rise to a man-like creature, resembling the higher anthropomorphous apes in so many respects. No doubt man, in comparison with most of his allies, has undergone an extraordinary amount of modification, chiefly in consequence of his greatly developed brain and erect position; nevertheless we should bear in mind that he “is but one of several exceptional forms of Primates.”[267]
Every naturalist, who believes in the principle of evolution, will grant that the two main divisions of the Simiadæ, namely the Catarhine and Platyrhine monkeys, with their sub-groups, have all proceeded from some one extremely ancient progenitor. The early descendants of this progenitor, before they had diverged to any considerable extent from each other, would still have formed a single natural group; but some of the species or incipient genera would have already begun to indicate by their diverging characters the future distinctive marks of the Catarhine and Platyrhine divisions. Hence the members of this supposed ancient group would not have been so uniform in their dentition or in the structure of their nostrils, as are the existing Catarhine monkeys in one way and the Platyrhines in another way, but would have resembled in this respect the allied Lemuridæ which differ greatly from each other in the form of their muzzles,[268] and to an extraordinary degree in their dentition.
The Catarhine and Platyrhine monkeys agree in a multitude of characters, as is shewn by their unquestionably belonging to one and the same Order. The many characters which they possess in common can hardly have been independently acquired by so many distinct species; so that these characters must have been inherited. But an ancient form which possessed many characters common to the Catarhine and Platyrhine monkeys, and others in an intermediate condition, and some few perhaps distinct from those now present in either group, would undoubtedly have been ranked, if seen by a naturalist, as an ape or monkey. And as man under a genealogical point of view belongs to the Catarhine or Old World stock, we must conclude, however much the conclusion may revolt our pride, that our early progenitors would have been properly thus designated.[269] But we must not fall into the error of supposing that the early progenitor of the whole Simian stock, including man, was identical with, or even closely resembled, any existing ape or monkey.
On the Birthplace and Antiquity of Man.—We are naturally led to enquire where was the birthplace of man at that stage of descent when our progenitors diverged from the Catarhine stock. The fact that they belonged to this stock clearly shews that they inhabited the Old World; but not Australia nor any oceanic island, as we may infer from the laws of geographical distribution. In each great region of the world the living mammals are closely related to the extinct species of the same region. It is therefore probable that Africa was formerly inhabited by extinct apes closely allied to the gorilla and chimpanzee; and as these two species are now man’s nearest allies, it is somewhat more probable that our early progenitors lived on the African continent than elsewhere. But it is useless to speculate on this subject, for an ape nearly as large as a man, namely the Dryopithecus of Lartet, which was closely allied to the anthropomorphous Hylobates, existed in Europe during the Upper Miocene period; and since so remote a period the earth has certainly undergone many great revolutions, and there has been ample time for migration on the largest scale.
At the period and place, whenever and wherever it may have been, when man first lost his hairy covering, he probably inhabited a hot country; and this would have been favourable for a frugiferous diet, on which, judging from analogy, he subsisted. We are far from knowing how long ago it was when man first diverged from the Catarhine stock; but this may have occurred at an epoch as remote as the Eocene period; for the higher apes had diverged from the lower apes as early as the Upper Miocene period, as shewn by the existence of the Dryopithecus. We are also quite ignorant at how rapid a rate organisms, whether high or low in the scale, may under favourable circumstances be modified: we know, however, that some have retained the same form during an enormous lapse of time. From what we see going on under domestication, we learn that within the same period some of the co-descendants of the same species may be not at all changed, some a little, and some greatly changed. Thus it may have been with man, who has undergone a great amount of modification in certain characters in comparison with the higher apes.
The great break in the organic chain between man and his nearest allies, which cannot be bridged over by any extinct or living species, has often been advanced as a grave objection to the belief that man is descended from some lower form; but this objection will not appear of much weight to those who, convinced by general reasons, believe in the general principle of evolution. Breaks incessantly occur in all parts of the series, some being wide, sharp and defined, others less so in various degrees; as between the orang and its nearest allies—between the Tarsius and the other Lemuridæ—between the elephant and in a more striking manner between the Ornithorhynchus or Echidna, and other mammals. But all these breaks depend merely on the number of related forms which have become extinct. At some future period, not very distant as measured by centuries, the civilised races of man will almost certainly exterminate and replace throughout the world the savage races. At the same time the anthropomorphous apes, as Professor Schaaffhausen has remarked,[270] will no doubt be exterminated. The break will then be rendered wider, for it will intervene between man in a more civilised state, as we may hope, than the Caucasian, and some ape as low as a baboon, instead of as at present between the negro or Australian and the gorilla.
With respect to the absence of fossil remains, serving to connect man with his ape-like progenitors, no one will lay much stress on this fact, who will read Sir C. Lyell’s discussion,[271] in which he shews that in all the vertebrate classes the discovery of fossil remains has been an extremely slow and fortuitous process. Nor should it be forgotten that those regions which are the most likely to afford remains connecting man with some extinct ape-like creature, have not as yet been searched by geologists.
Lower Stages in the Genealogy of Man.—We have seen that man appears to have diverged from the Catarhine or Old World division of the Simiadæ, after these had diverged from the New World division. We will now endeavour to follow the more remote traces of his genealogy, trusting in the first place to the mutual affinities between the various classes and orders, with some slight aid from the periods, as far as ascertained, of their successive appearance on the earth. The Lemuridæ stand below and close to the Simiadæ, constituting a very distinct family of the Primates, or, according to Häckel, a distinct Order. This group is diversified and broken to an extraordinary degree, and includes many aberrant forms. It has, therefore, probably suffered much extinction. Most of the remnants survive on islands, namely in Madagascar and in the islands of the Malayan archipelago, where they have not been exposed to such severe competition as they would have been on well-stocked continents. This group likewise presents many gradations, leading, as Huxley remarks,[272] “insensibly from the crown and summit of the animal creation down to creatures from which there is but a step, as it seems, to the lowest, smallest, and least intelligent of the placental mammalia.” From these various considerations it is probable that the Simiadæ were originally developed from the progenitors of the existing Lemuridæ; and these in their turn from forms standing very low in the mammalian series.