Now let us apply these generally-admitted principles to the races of man, viewing him in the same spirit as a naturalist would any other animal. In regard to the amount of difference between the races, we must make some allowance for our nice powers of discrimination gained by the long habit of observing ourselves. In India, as Elphinstone remarks,[282] although a newly-arrived European cannot at first distinguish the various native races, yet they soon appear to him extremely dissimilar; and the Hindoo cannot at first perceive any difference between the several European nations. Even the most distinct races of man, with the exception of certain negro tribes, are much more like each other in form than would at first be supposed. This is well shewn by the French photographs in the Collection Anthropologique du Muséum of the men belonging to various races, the greater number of which, as many persons to whom I have shown them have remarked, might pass for Europeans. Nevertheless, these men if seen alive would undoubtedly appear very distinct, so that we are clearly much influenced in our judgment by the mere colour of the skin and hair, by slight differences in the features, and by expression.

There is, however, no doubt that the various races, when carefully compared and measured, differ much from each other,—as in the texture of the hair, the relative proportions of all parts of the body,[283] the capacity of the lungs, the form and capacity of the skull, and even in the convolutions of the brain.[284] But it would be an endless task to specify the numerous points of structural difference. The races differ also in constitution, in acclimatisation, and in liability to certain diseases. Their mental characteristics are likewise very distinct; chiefly as it would appear in their emotional, but partly in their intellectual, faculties. Every one who has had the opportunity of comparison, must have been struck with the contrast between the taciturn, even morose, aborigines of S. America and the light-hearted, talkative negroes. There is a nearly similar contrast between the Malays and the Papuans,[285] who live under the same physical conditions, and are separated from each other only by a narrow space of sea.

We will first consider the arguments which may be advanced in favour of classing the races of man as distinct species, and then those on the other side. If a naturalist, who had never before seen such beings, were to compare a Negro, Hottentot, Australian, or Mongolian, he would at once perceive that they differed in a multitude of characters, some of slight and some of considerable importance. On inquiry he would find that they were adapted to live under widely different climates, and that they differed somewhat in bodily constitution and mental disposition. If he were then told that hundreds of similar specimens could be brought from the same countries, he would assuredly declare that they were as good species as many to which he had been in the habit of affixing specific names. This conclusion would be greatly strengthened as soon as he had ascertained that these forms had all retained the same character for many centuries; and that negroes, apparently identical with existing negroes, had lived at least 4000 years ago.[286] He would also hear from an excellent observer, Dr. Lund,[287] that the human skulls found in the caves of Brazil, entombed with many extinct mammals, belonged to the same type as that now prevailing throughout the American Continent.

Our naturalist would then perhaps turn to geographical distribution, and he would probably declare that forms differing not only in appearance, but fitted for the hottest and dampest or driest countries, as well as for the arctic regions, must be distinct species. He might appeal to the fact that no one species in the group next to man, namely the Quadrumana, can resist a low temperature or any considerable change of climate; and that those species which come nearest to man have never been reared to maturity, even under the temperate climate of Europe. He would be deeply impressed with the fact, first noticed by Agassiz,[288] that the different races of man are distributed over the world in the same zoological provinces, as those inhabited by undoubtedly distinct species and genera of mammals. This is manifestly the case with the Australian, Mongolian, and Negro races of man; in a less well-marked manner with the Hottentots; but plainly with the Papuans and Malays, who are separated, as Mr. Wallace has shewn, by nearly the same line which divides the great Malayan and Australian zoological provinces. The aborigines of America range throughout the Continent; and this at first appears opposed to the above rule, for most of the productions of the Southern and Northern halves differ widely; yet some few living forms, as the opossum, range from the one into the other, as did formerly some of the gigantic Edentata. The Esquimaux, like other Arctic animals, extend round the whole polar regions. It should be observed that the mammalian forms which inhabit the several zoological provinces, do not differ from each other in the same degree; so that it can hardly be considered as an anomaly that the Negro differs more, and the American much less, from the other races of man than do the mammals of the same continents from those of the other provinces. Man, it may be added, does not appear to have aboriginally inhabited any oceanic island; and in this respect he resembles the other members of his class.

In determining whether the varieties of the same kind of domestic animal should be ranked as specifically distinct, that is, whether any of them are descended from distinct wild species, every naturalist would lay much stress on the fact, if established, of their external parasites being specifically distinct. All the more stress would be laid on this fact, as it would be an exceptional one, for I am informed by Mr. Denny that the most different kinds of dogs, fowls, and pigeons, in England, are infested by the same species of Pediculi or lice. Now Mr. A. Murray has carefully examined the Pediculi collected in different countries from the different races of man;[289] and he finds that they differ, not only in colour, but in the structure of their claws and limbs. In every case in which numerous specimens were obtained the differences were constant. The surgeon of a whaling ship in the Pacific assured me that when the Pediculi, with which some Sandwich Islanders on board swarmed, strayed on to the bodies of the English sailors, they died in the course of three or four days. These Pediculi were darker coloured and appeared different from those proper to the natives of Chiloe in South America, of which he gave me specimens. These, again, appeared larger and much softer than European lice. Mr. Murray procured four kinds from Africa, namely from the Negroes of the Eastern and Western coasts, from the Hottentots and Caffres; two kinds from the natives of Australia; two from North, and two from South America. In these latter cases it may be presumed that the Pediculi came from natives inhabiting different districts. With insects slight structural differences, if constant, are generally esteemed of specific value: and the fact of the races of man being infested by parasites, which appear to be specifically distinct, might fairly be urged as an argument that the races themselves ought to be classed as distinct species.

Our supposed naturalist having proceeded thus far in his investigation, would next inquire whether the races of men, when crossed, were in any degree sterile. He might consult the work[290] of a cautious and philosophical observer, Professor Broca; and in this he would find good, evidence that some races were quite fertile together; but evidence of an opposite nature in regard to other races. Thus it has been asserted that the native women of Australia and Tasmania rarely produce children to European men; the evidence, however, on this head has now been shewn to be almost valueless. The half-castes are killed by the pure blacks; and an account has lately been published of eleven half-caste youths murdered and burnt at the same time, whose remains were found by the police.[291] Again, it has often been said that when mulattoes intermarry they produce few children; on the other hand, Dr. Bachman of Charlestown[292] positively asserts that he has known mulatto families which have intermarried for several generations, and have continued on an average as fertile as either pure whites or pure blacks. Inquiries formerly made by Sir C. Lyell on this subject led him, as he informs me, to the same conclusion. In the United States the census for the year 1854 included, according to Dr. Bachman, 405,751 mulattoes; and this number, considering all the circumstances of the case, seems small; but it may partly be accounted for by the degraded and anomalous position of the class, and by the profligacy of the women. A certain amount of absorption of mulattoes into negroes must always be in progress; and this would lead to an apparent diminution of the former. The inferior vitality of mulattoes is spoken of in a trustworthy work[293] as a well-known phenomenon; but this is a different consideration from their lessened fertility; and can hardly be advanced as a proof of the specific distinctness of the parent races. No doubt both animal and vegetable hybrids, when produced from extremely distinct species, are liable to premature death; but the parents of mulattoes cannot be put under the category of extremely distinct species. The common Mule, so notorious for long life and vigour, and yet so sterile, shews how little necessary connection there is in hybrids between lessened fertility and vitality: other analogous cases could be added.

Even if it should hereafter be proved that all the races of men were perfectly fertile together, he who was inclined from other reasons to rank them as distinct species, might with justice argue that fertility and sterility are not safe criterions of specific distinctness. We know that these qualities are easily affected by changed conditions of life or by close inter-breeding, and that they are governed by highly complex laws, for instance that of the unequal fertility of reciprocal crosses between the same two species. With forms which must be ranked as undoubted species, a perfect series exists from those which are absolutely sterile when crossed, to those which are almost or quite fertile. The degrees of sterility do not coincide strictly with the degrees of difference in external structure or habits of life. Man in many respects may be compared with those animals which have long been domesticated, and a large body of evidence can be advanced in favour of the Pallasian doctrine[294] that domestication tends to eliminate the sterility which is so general a result of the crossing of species in a state of nature. From these several considerations, it may be justly urged that the perfect fertility of the intercrossed races of man, if established, would not absolutely preclude us from ranking them as distinct species.

Independently of fertility, the character of the offspring from a cross has sometimes been thought to afford evidence whether the parent-forms ought to be ranked as species or varieties; but after carefully studying the evidence, I have come to the conclusion that no general rules of this kind can be trusted. Thus with mankind the offspring of distinct races resemble in all respects the offspring of true species and of varieties. This is shewn, for instance, by the manner in which the characters of both parents are blended, and by one form absorbing another through repeated crosses. In this latter case the progeny both of crossed species and varieties retain for a long period a tendency to revert to their ancestors, especially to that one which is prepotent in transmission. When any character has suddenly appeared in a race or species as the result of a single act of variation, as is general with monstrosities,[295] and this race is crossed with another not thus characterised, the characters in question do not commonly appear in a blended condition in the young, but are transmitted to them either perfectly developed or not at all. As with the crossed races of man cases of this kind rarely or never occur, this may be used as an argument against the view suggested by some ethnologists, namely that certain characters, for instance the blackness of the negro, first appeared as a sudden variation or sport. Had this occurred, it is probable that mulattoes would often have been born, either completely black or completely white.

We have now seen that a naturalist might feel himself fully justified in ranking the races of man as distinct species; for he has found that they are distinguished by many differences in structure and constitution, some being of importance. These differences have, also, remained nearly constant for very long periods of time. He will have been in some degree influenced by the enormous range of man, which is a great anomaly in the class of mammals, if mankind be viewed as a single species. He will have been struck with the distribution of the several so-called races, in accordance with that of other undoubtedly distinct species of mammals. Finally he might urge that the mutual fertility of all the races has not as yet been fully proved; and even if proved would not be an absolute proof of their specific identity.

On the other side of the question, if our supposed naturalist were to enquire whether the forms of man kept distinct like ordinary species, when mingled together in large numbers in the same country, he would immediately discover that this was by no means the case. In Brazil he would behold an immense mongrel population of Negroes and Portuguese; in Chiloe and other parts of South America, he would behold the whole population consisting of Indians and Spaniards blended in various degrees.[296] In many parts of the same continent he would meet with the most complex crosses between Negroes, Indians, and Europeans; and such triple crosses afford the severest test, judging from the vegetable kingdom, of the mutual fertility of the parent-forms. In one island of the Pacific he would find a small population of mingled Polynesian and English blood; and in the Viti Archipelago a population of Polynesians and Negritos crossed in all degrees. Many analogous cases could be added, for instance, in South Africa. Hence the races of man are not sufficiently distinct to co-exist without fusion; and this it is, which in all ordinary cases affords the usual test of specific distinctness.