Turning to primeval times when men had only doubtfully attained the rank of manhood, they would probably have lived, as already stated, either as polygamists or temporarily as monogamists. Their intercourse, judging from analogy, would not then have been promiscuous. They would, no doubt, have defended their females to the best of their power from enemies of all kinds, and would probably have hunted for their subsistence, as well as for that of their offspring. The most powerful and able males would have succeeded best in the struggle for life and in obtaining attractive females. At this early period the progenitors of man, from having only feeble powers of reason, would not have looked forward to distant contingencies. They would have been governed more by their instincts and even less by their reason than are savages at the present day. They would not at that period have partially lost one of the strongest of all instincts, common to all the lower animals, namely the love of their young offspring; and consequently they would not have practised infanticide. There would have been no artificial scarcity of women, and polyandry would not have been followed; there would have been no early betrothals; women would not have been valued as mere slaves; both sexes, if the females as well as the males were permitted to exert any choice, would have chosen their partners, not for mental charms, or property, or social position, but almost solely from external appearance. All the adults would have married or paired, and all the offspring, as far as that was possible, would have been reared; so that the struggle for existence would have been periodically severe to an extreme degree. Thus during these primordial times all the conditions for sexual selection would have been much more favourable than at a later period, when man had advanced in his intellectual powers, but had retrograded in his instincts. Therefore, whatever influence sexual selection may have had in producing the differences between the races of man, and between man and the higher Quadrumana, this influence would have been much more powerful at a very remote period than at the present day.

On the Manner of Action of Sexual Selection with mankind.—With primeval men under the favourable conditions just stated, and with those savages who at the present time enter into any marriage tie (but subject to greater or less interference according as the habits of female infanticide, early betrothals, &c., are more or less practised), sexual selection will probably have acted in the following manner. The strongest and most vigorous men,—those who could best defend and hunt for their families, and during later times the chiefs or head-men,—those who were provided with the best weapons and who possessed the most property, such as a larger number of dogs or other animals, would have succeeded in rearing a greater average number of offspring, than would the weaker, poorer and lower members of the same tribes. There can, also, be no doubt that such men would generally have been able to select the more attractive women. At present the chiefs of nearly every tribe throughout the world succeed in obtaining more than one wife. Until recently, as I hear from Mr. Mantell, almost every girl in New Zealand, who was pretty, or promised to be pretty, was tapu to some chief. With the Kafirs, as Mr. C. Hamilton states,[462] “the chiefs generally have the pick of the women for many miles round, and are most persevering in establishing or confirming their privilege.” We have seen that each race has its own style of beauty, and we know that it is natural to man to admire each characteristic point in his domestic animals, dress, ornaments, and personal appearance, when carried a little beyond the common standard. If then the several foregoing propositions be admitted, and I cannot see that they are doubtful, it would be an inexplicable circumstance, if the selection of the more attractive women by the more powerful men of each tribe, who would rear on an average a greater number of children, did not after the lapse of many generations modify to a certain extent the character of the tribe.

With our domestic animals, when a foreign breed is introduced into a new country, or when a native breed is long and carefully attended to, either for use or ornament, it is found after several generations to have undergone, whenever the means of comparison exist, a greater or less amount of change. This follows from unconscious selection during a long series of generations—that is, the preservation of the most approved individuals—without any wish or expectation of such a result on the part of the breeder. So again, if two careful breeders rear during many years animals of the same family, and do not compare them together or with a common standard, the animals are found after a time to have become to the surprise of their owners slightly different.[463] Each breeder has impressed, as Von Nathusius well expresses it, the character of his own mind—his own taste and judgment—on his animals. What reason, then, can be assigned why similar results should not follow from the long-continued selection of the most admired women by those men of each tribe, who were able to rear to maturity the greater number of children? This would be unconscious selection, for an effect would be produced, independently of any wish or expectation on the part of the men who preferred certain women to others.

Let us suppose the members of a tribe, in which some form of marriage was practised, to spread over an unoccupied continent; they would soon split up into distinct hordes, which would be separated from each other by various barriers, and still more effectually by the incessant wars between all barbarous nations. The hordes would thus be exposed to slightly different conditions and habits of life, and would sooner or later come to differ in some small degree. As soon as this occurred, each isolated tribe would form for itself a slightly different standard of beauty;[464] and then unconscious selection would come into action through the more powerful and leading savages preferring certain women to others. Thus the differences between the tribes, at first very slight, would gradually and inevitably be increased to a greater and greater degree.

With animals in a state of nature, many characters proper to the males, such as size, strength, special weapons, courage and pugnacity, have been acquired through the law of battle. The semi-human progenitors of man, like their allies the Quadrumana, will almost certainly have been thus modified; and, as savages still fight for the possession of their women, a similar process of selection has probably gone on in a greater or less degree to the present day. Other characters proper to the males of the lower animals, such as bright colours and various ornaments, have been acquired by the more attractive males having been preferred by the females. There are, however, exceptional cases in which the males, instead of having been the selected, have been the selectors. We recognise such cases by the females having been rendered more highly ornamented than the males,—their ornamental characters having been transmitted exclusively or chiefly to their female offspring. One such case has been described in the order to which man belongs, namely, with the Rhesus monkey.

Man is more powerful in body and mind than woman, and in the savage state he keeps her in a far more abject state of bondage than does the male of any other animal; therefore it is not surprising that he should have gained the power of selection. Women are everywhere conscious of the value of their beauty; and when they have the means, they take more delight in decorating themselves with all sorts of ornaments than do men. They borrow the plumes of male birds, with which nature decked this sex in order to charm the females. As women have long been selected for beauty, it is not surprising that some of the successive variations should have been transmitted in a limited manner; and consequently that women should have transmitted their beauty in a somewhat higher degree to their female than to their male offspring. Hence women have become more beautiful, as most persons will admit, than men. Women, however, certainly transmit most of their characters, including beauty, to their offspring of both sexes; so that the continued preference by the men of each race of the more attractive women, according to their standard of taste, would tend to modify in the same manner all the individuals of both sexes belonging to the race.

With respect to the other form of sexual selection (which with the lower animals is much the most common), namely, when the females are the selectors, and accept only those males which excite or charm them most, we have reason to believe that it formerly acted on the progenitors of man. Man in all probability owes his beard, and perhaps some other characters, to inheritance from an ancient progenitor who gained in this manner his ornaments. But this form of selection may have occasionally acted during later times; for in utterly barbarous tribes the women have more power in choosing, rejecting, and tempting their lovers, or of afterwards changing their husbands, than might have been expected. As this is a point of some importance, I will give in detail such evidence as I have been able to collect.

Hearne describes how a woman in one of the tribes of Arctic America repeatedly ran away from her husband and joined a beloved man; and with the Charruas of S. America, as Azara states, the power of divorce is perfectly free. With the Abipones, when a man chooses a wife he bargains with the parents about the price. But “it frequently happens that the girl rescinds what has been agreed upon between the parents and the bridegroom, obstinately rejecting the very mention of marriage.” She often runs away, hides herself, and thus eludes the bridegroom. In the Fiji Islands the man seizes on the woman whom he wishes for his wife by actual or pretended force; but “on reaching the home of her abductor, should she not approve of the match, she runs to some one who can protect her; if, however, she is satisfied, the matter is settled forthwith.” In Tierra del Fuego a young man first obtains the consent of the parents by doing them some service, and then he attempts to carry off the girl; “but if she is unwilling, she hides herself in the woods until her admirer is heartily tired of looking for her, and gives up the pursuit; but this seldom happens.” With the Kalmucks there is a regular race between the bride and bridegroom, the former having a fair start; and Clarke “was assured that no instance occurs of a girl being caught, unless she has a partiality to the pursuer.” So with the wild tribes of the Malay archipelago there is a similar racing match; and it appears from M. Bourien’s account, as Sir J. Lubbock remarks, that “the race ‘is not to the swift, nor the battle to the strong,’ but to the young man who has the good fortune to please his intended bride.”

Turning to Africa: the Kafirs buy their wives, and girls are severely beaten by their fathers if they will not accept a chosen husband; yet it is manifest from many facts given by the Rev. Mr. Shooter, that they have considerable power of choice. Thus very ugly, though rich men, have been known to fail in getting wives. The girls, before consenting to be betrothed, compel the men to shew themselves off, first in front and then behind, and “exhibit their paces.” They have been known to propose to a man, and they not rarely run away with a favoured lover. With the degraded bushwomen of S. Africa, “when a girl has grown up to womanhood without having been betrothed, which, however, does not often happen, her lover must gain her approbation, as well as that of the parents.”[465] Mr. Winwood Reade made inquiries for me with respect to the negroes of Western Africa, and he informs me that “the women, at least among the more intelligent Pagan tribes, have no difficulty in getting the husbands whom they may desire, although it is considered unwomanly to ask a man to marry them. They are quite capable of falling in love, and of forming tender, passionate, and faithful attachments.”

We thus see that with savages the women are not in quite so abject a state in relation to marriage as has often been supposed. They can tempt the men whom they prefer, and can sometimes reject those whom they dislike, either before or after marriage. Preference on the part of the women, steadily acting in any one direction, would ultimately affect the character of the tribe; for the women would generally choose not merely the handsomer men, according to their standard of taste, but those who were at the same time best able to defend and support them. Such well-endowed pairs would commonly rear a larger number of offspring than the less well endowed. The same result would obviously follow in a still more marked manner if there was selection on both sides; that is if the more attractive, and at the same time more powerful men were to prefer, and were preferred by, the more attractive women. And these two forms of selection seem actually to have occurred, whether or not simultaneously, with mankind, especially during the earlier periods of our long history.