If it could be shown that worms generally excavate their burrows at right angles to an inclined surface, and this would be their shortest course for bringing up earth from beneath, then as the old burrows collapsed from the weight of the superincumbent soil, the collapsing would inevitably cause the whole bed of vegetable mould to sink or slide slowly down the inclined surface. But to ascertain the direction of many burrows was found too difficult and troublesome. A straight piece of wire was, however, pushed into twenty-five burrows on several sloping fields, and in eight cases the burrows were nearly at right angles to the slope; whilst in the remaining cases they were indifferently directed at various angles, either upwards or downwards with respect to the slope.
In countries where the rain is very heavy, as in the tropics, the castings appear, as might have been expected, to be washed down in a greater degree than in England. Mr. Scott informs me that near Calcutta the tall columnar castings (previously described), the diameter of which is usually between 1 and 1½ inch, subside on a level surface, after heavy rain, into almost circular, thin, flat discs, between 3 and 4 and sometimes 5 inches in diameter. Three fresh castings, which had been ejected in the Botanic Gardens “on a slightly inclined, grass-covered, artificial bank of loamy clay,” were carefully measured, and had a mean height of 2.17, and a mean diameter of 1.43 inches; these after heavy rain, formed elongated patches of earth, with a mean length in the direction of the slope of 5.83 inches. As the earth had spread very little up the slope, a large part, judging from the original diameter of these castings, must have flowed bodily downwards about 4 inches. Moreover some of the finest earth of which they were composed must have been washed completely away to a still greater distance. In drier sites near Calcutta, a species of worm ejects its castings, not in vermiform masses, but in little pellets of varying sizes: these are very numerous in some places, and Mr. Scott says that they “are washed away by every shower.”
I was led to believe that a considerable quantity of fine earth is washed quite away from castings during rain, from the surfaces of old ones being often studded with coarse particles. Accordingly a little fine precipitated chalk, moistened with saliva or gum-water, so as to be slightly viscid and of the same consistence as a fresh casting, was placed on the summits of several castings and gently mixed with them. These castings were then watered through a very fine rose, the drops from which were closer together than those of rain, but not nearly so large as those in a thunderstorm; nor did they strike the ground with nearly so much force as drops during heavy rain. A casting thus treated subsided with surprising slowness, owing as I suppose to its viscidity. It did not flow bodily down the grass-covered surface of the lawn, which was here inclined at an angle of 16° 20′; nevertheless many particles of the chalk were found three inches below the casting. The experiment was repeated on three other castings on different parts of the lawn, which sloped at 2° 30′, 3° and 6°; and particles of chalk could be seen between 4 and 5 inches below the casting; and after the surface had become dry, particles were found in two cases at a distance of 5 and 6 inches. Several other castings with precipitated chalk placed on their summits were left to the natural action of the rain. In one case, after rain which was not heavy, the casting was longitudinally streaked with white. In two other cases the surface of the ground was rendered somewhat white for a distance of one inch from the casting; and some soil collected at a distance of 2½ inches, where the slope was 7°, effervesced slightly when placed in acid. After one or two weeks, the chalk was wholly or almost wholly washed away from all the castings on which it had been placed, and these had recovered their natural colour.
It may be here remarked that after very heavy rain shallow pools may be seen on level or nearly level fields, where the soil is not very porous, and the water in them is often slightly muddy; when such little pools have dried, the leaves and blades of grass at their bottoms are generally coated with a thin layer of mud. This mud I believe is derived in large part from recently ejected castings.
Dr. King informs me that the majority of the before described gigantic castings, which he found on a fully exposed, bare, gravelly knoll on the Nilgiri Mountains in India, had been more or less weathered by the previous north-east monsoon; and most of them presented a subsided appearance. The worms here eject their castings only during the rainy season; and at the time of Dr. King’s visit no rain had fallen for 110 days. He carefully examined the ground between the place where these huge castings lay, and a little watercourse at the base of the knoll, and nowhere was there any accumulation of fine earth, such as would necessarily have been left by the disintegration of the castings if they had not been wholly removed. He therefore has no hesitation in asserting that the whole of these huge castings are annually washed during the two monsoons (when about 100 inches of rain fall) into the little water-course, and thence into the plains lying below at a depth of 3000 or 4000 feet.
Castings ejected before or during dry weather become hard, sometimes surprisingly hard, from the particles of earth having been cemented together by the intestinal secretions. Frost seems to be less effective in their disintegration than might have been expected. Nevertheless they readily disintegrate into small pellets, after being alternately moistened with rain and again dried. Those which have flowed during rain down a slope, disintegrate in the same manner. Such pellets often roll a little down any sloping surface; their descent being sometimes much aided by the wind. The whole bottom of a broad dry ditch in my grounds, where there were very few fresh castings, was completely covered with these pellets or disintegrated castings, which had rolled down the steep sides, inclined at an angle of 27°.
Near Nice, in places where the great cylindrical castings, previously described, abound, the soil consists of very fine arenaceo-calcareous loam; and Dr. King informs me that these castings are extremely liable to crumble during dry weather into small fragments, which are soon acted on by rain, and then sink down so as to be no longer distinguishable from the surrounding soil. He sent me a mass of such disintegrated castings, collected on the top of a bank, where none could have rolled down from above. They must have been ejected within the previous five or six months, but they now consisted of more or less rounded fragments of all sizes, from ¾ of an inch in diameter to minute grains and mere dust. Dr. King witnessed the crumbling process whilst drying some perfect castings, which he afterwards sent me. Mr. Scott also remarks on the crumbling of the castings near Calcutta and on the mountains of Sikkim during the hot and dry season.
When the castings near Nice had been ejected on an inclined surface, the disintegrated fragments rolled downwards, without losing their distinctive shape; and in some places could “be collected in basketfuls.” Dr. King observed a striking instance of this fact on the Corniche road, where a drain, about 2½ feet wide and 9 inches deep, had been made to catch the surface drainage from the adjoining hill-side. The bottom of this drain was covered for a distance of several hundred yards, to a depth of from 1½ to 3 inches, by a layer of broken castings, still retaining their characteristic shape. Nearly all these innumerable fragments had rolled down from above, for extremely few castings had been ejected in the drain itself. The hill-side was steep, but varied much in inclination, which Dr. King estimated at from 30° to 60° with the horizon. He climbed up the slope, and “found every here and there little embankments, formed by fragments of the castings that had been arrested in their downward progress by irregularities of the surface, by stones, twigs, &c. One little group of plants of Anemone hortensis had acted in this manner, and quite a small bank of soil had collected round it. Much of this soil had crumbled down, but a great deal of it still retained the form of castings.” Dr. King dug up this plant, and was struck with the thickness of the soil which must have recently accumulated over the crown of the rhizoma, as shown by the length of the bleached petioles, in comparison with those of other plants of the same kind, where there had been no such accumulation. The earth thus accumulated had no doubt been secured (as I have everywhere seen) by the smaller roots of the plants. After describing this and other analogous cases, Dr. King concludes: “I can have no doubt that worms help greatly in the process of denudation.”
Ledges of earth on steep hill-sides.—Little horizontal ledges, one above another, have been observed on steep grassy slopes in many parts of the world. The formation has been attributed to animals travelling repeatedly along the slope in the same horizontal lines while grazing, and that they do thus move and use the ledges is certain; but Professor Henslow (a most careful observer) told Sir J. Hooker that he was convinced that this was not the sole cause of their formation. Sir J. Hooker saw such ledges on the Himalayan and Atlas ranges, where there were no domesticated animals and not many wild ones; but these latter would, it is probable, use the ledges at night while grazing like our domesticated animals. A friend observed for me the ledges on the Alps of Switzerland, and states that they ran at 3 or 4 ft. one above the other, and were about a foot in breadth. They had been deeply pitted by the feet of grazing cows. Similar ledges were observed by the same friend on our Chalk downs, and on an old talus of chalk-fragments (thrown out of a quarry) which had become clothed with turf.
My son Francis examined a Chalk escarpment near Lewes; and here on a part which was very steep, sloping at 40° with the horizon, about 30 flat ledges extended horizontally for more than 100 yards, at an average distance of about 20 inches, one beneath the other. They were from 9 to 10 inches in breadth. When viewed from a distance they presented a striking appearance, owing to their parallelism; but when examined closely, they were seen to be somewhat sinuous, and one often ran into another, giving the appearance of the ledge having forked into two. They are formed of light-coloured earth, which on the outside, where thickest, was in one case 9 inches, and in another case between 6 and 7 inches in thickness. Above the ledges, the thickness of the earth over the chalk was in the former case 4 and in the latter only 3 inches. The grass grew more vigorously on the outer edges of the ledges than on any other part of the slope, and here formed a tufted fringe. Their middle part was bare, but whether this had been caused by the trampling of sheep, which sometimes frequent the ledges, my son could not ascertain. Nor could he feel sure how much of the earth on the middle and bare parts, consisted of disintegrated worm-castings which had rolled down from above; but he felt convinced that some had thus originated; and it was manifest that the ledges with their grass-fringed edges would arrest any small object rolling down from above.