My reasons have now been assigned for believing that specific forms are not immutable creations[{518}]. The terms used by naturalists of affinity, unity of type, adaptive characters, the metamorphosis and abortion of organs, cease to be metaphorical expressions and become intelligible facts. We no longer look at an organic being as a savage does at a ship[{519}] or other great work of art, as at a thing wholly beyond his comprehension, but as a production that has a history which we may search into. How interesting do all instincts become when we speculate on their origin as hereditary habits, or as slight congenital modifications of former instincts perpetuated by the individuals so characterised having been preserved. When we look at every complex instinct and mechanism as the summing up of a long history of contrivances, each most useful to its possessor, nearly in the same way as when we look at a great mechanical invention as the summing up of the labour, the experience, the reason, and even the blunders of numerous workmen. How interesting does the geographical distribution of all organic beings, past and present, become as throwing light on the ancient geography of the world. Geology loses glory[{520}] from the imperfection of its archives, but it gains in the immensity of its subject. There is much grandeur in looking at every existing organic being either as the lineal successor of some form now buried under thousands of feet of solid rock, or as being the co-descendant of that buried form of some more ancient and utterly lost inhabitant of this world. It accords with what we know of the laws impressed by the Creator[{521}] on matter that the production and extinction of forms should, like the birth and death of individuals, be the result of secondary means. It is derogatory that the Creator of countless Universes should have made by individual acts of His will the myriads of creeping parasites and worms, which since the earliest dawn of life have swarmed over the land and in the depths of the ocean. We cease to be astonished[{522}] that a group of animals should have been formed to lay their eggs in the bowels and flesh of other sensitive beings; that some animals should live by and even delight in cruelty; that animals should be led away by false instincts; that annually there should be an incalculable waste of the pollen, eggs and immature beings; for we see in all this the inevitable consequences of one great law, of the multiplication of organic beings not created immutable. From death, famine, and the struggle for existence, we see that the most exalted end which we are capable of conceiving, namely, the creation of the higher animals[{523}], has directly proceeded. Doubtless, our first impression is to disbelieve that any secondary law could produce infinitely numerous organic beings, each characterised by the most exquisite workmanship and widely extended adaptations: it at first accords better with our faculties to suppose that each required the fiat of a Creator. There[{524}] is a [simple] grandeur in this view of life with its several powers of growth, reproduction and of sensation, having been originally breathed into matter under a few forms, perhaps into only one[{525}], and that whilst this planet has gone cycling onwards according to the fixed laws of gravity and whilst land and water have gone on replacing each other—that from so simple an origin, through the selection of infinitesimal varieties, endless forms most beautiful and most wonderful have been evolved.
INDEX
For the names of Authors, Birds, Mammals (including names of classes) and Plants, see sub-indexes under Authors, Birds, Mammals and Plants.
- Acquired characters, see Characters
- Affinities and classification, [35]
- America, fossils, [177]
- Analogy, resemblance by, [36], [82], [199], [205], [211]
- Animals, marine, preservation of as fossils, [25], [139], [141];
- Australia, fossils, [177]
- Authors, Names of:—
- Ackerman on hybrids, [11];
- Bakewell, [9], [91];
- Bateson, W., [xxix], [69] n., [217];
- Bellinghausen, [124];
- Boitard and Corbié, [106] n.;
- Brougham, Lord, [17], [117];
- Brown, R., [233];
- Buckland on fossils, [24], [137], [145] n.;
- Buffon on woodpecker, [6];
- Bunbury (Sir H.), rules for selection, [67];
- Butler, S., [116] n.;
- d’Archiac, [146] n.;
- Darwin, C., origin of his evolutionary views, [xi]-[xv];
- Darwin, Mrs, letter to, [xxvi];
- Darwin, F., on Knight’s Law, [70] n.;
- Darwin, R. W., fact supplied by, [42] n., [223];
- Darwin and Wallace, joint paper by, [xxiv], [42] n.;
- De Candolle, [7], [47], [87], [204], [238];
- D’Orbigny, [124], [179] n.;
- Ehrenberg, [146] n.;
- Ewart on telegony, [108] n.;
- Falconer, [167];
- Forbes, E., [xxvii], [30], [146] n., [163] n., [165] n.;
- Gadow, Dr, [xxix];
- Gärtner, [98], [107];
- Goebel on Knight’s Law, [70] n.;
- Gould on distribution, [156];
- Gray, Asa, letter to, publication of in Linnean paper explained, [xxiv];
- Henslow, G., on evolution without selection, [63] n.;
- Henslow, J. S., [xxvii];
- Herbert on hybrids, [12], [98];
- —sterility of crocus, [99] n.;
- Hering, [116] n.;
- Hogg, [115] n.;
- Holland, Dr, [223];
- Hooker, J. D., [xxvii], [xxviii], [153] n.;
- Huber, P., [118];
- Hudson on woodpecker, [131] n.;
- Humboldt, [71], [166];
- Hunter, W., [114];
- Hutton, [27], [138];
- Huxley, [134] n.;
- Judd, [xi], [xiii], [xxix], [28], [141] n.;
- Knight, A., [3] n., [65], [xi];
- —on Domestication, [77];
- Knight-Darwin Law, [70] n.;
- Kölreuter, [12], [97], [98], [104], [232];
- Lamarck, [42] n., [47], [82], [146], [200];
- —reasons for his belief in mutability, [197];
- Lindley, [101];
- Linnean Society, joint paper, see Darwin and Wallace;
- Linnæus on sterility of Alpine plants, [101];
- —on generic characters, [201];
- Lonsdale, [145] n.;
- Lyell, [xxvii], [134] n., [138], [141] and n., [146] n., [159], [171], [173], [178];
- Macculloch, [124] n.;
- Macleay, W. S., [202];
- Magendie, [117];
- Malthus, [xv], [7], [88], [90];
- Marr, Dr, [xxix];
- Marshall, [65];
- Mivart, criticisms, [128] n.;
- Mozart as a child, his skill on the piano compared to instinct, [19] n.;
- Müller on consensual movements, [113];
- Murchison, [145] n.;
- Newton, Alfred, [132] n.;
- Owen, R., [xxvii], [219];
- Pallas, [68], [69];
- Pennant, [93] n.;
- Pliny on selection, [67];
- Poeppig, [113] n.;
- Prain, Col., [xxix];
- Rengger, sterility, [100];
- Richardson, [132] n.;
- Rutherford, H. W., [xxix];
- St Hilaire on races of dogs, [106];
- Smith, Jordan, [140];
- Sprengel, [233];
- Stapf, Dr, [xxix];
- Strickland, [xxvii];
- Suchetet, [97] n.;
- Thiselton-Dyer, Sir W., [xxix], [167];
- Wallace, [xxiv], [xxix], [30], [170] n.;
- Waterhouse, [125], [126];
- Western, Lord, [9], [65], [91];
- Whewell, [xxviii], [200];
- Woodward, H. B., [145] n.;
- Wrangel, [119] n.;
- Zacharias, Darwin’s letter to, [xv]
- Barriers and distribution, [30], [154], [157], [178]
- Bees, [113], [117];
- Beetles, abortive wings of, [45]
- Birds, transporting seeds, [169];
- Apteryx, [45], [236];
- Duck, [46], [61], [65], [128], [224] n.;
- Fowl, domestic, [59], [82] n., [97], [113], [114], [217];
- Goose, [72];
- —periodic habit, [124] n.;
- Grouse, hybridised, [97], [102];
- Guinea-fowl, [79];
- Hawk, sterility, [79];
- —periodic habit, [124];
- Opetiorynchus, [83];
- Orpheus, [31];
- Ostrich, distribution of, [158];
- Owl, white barn, [82];
- Partridge, infertility of, [102];
- Peacock, [79], [97], [102];
- Penguin, [128] n., [237];
- Petrel, [128] n.;
- Pheasant, [97], [102];
- Pigeon, [66], [82], [110] n., [113], [114], [116], [117], [129], [135]; see Wood-pigeon;
- Rhea, [158];
- Robins, increase in numbers, [88], [90];
- Rock-thrush of Guiana, [93];
- Swan, species of, [105];
- Tailor-bird, [18], [118];
- Turkey, Australian bush-turkey, [121] n., [122];
- Tyrannus, [31];
- Water-ouzel, [18] n., [120];
- Woodcock, loss of migratory instinct, [120];
- Woodpecker, [6], [16], [128] n., [148];
- Wood-pigeon, [122];
- Wren, gold-crested, [120];
- Birds, Names of:—
- Breeds, domestic, parentage of, [71]
- Brothers, death of by same peculiar disease in old age, [42] n., [44] n., [223]
- Bud variation, [58]; see Sports
- Butterfly, cabbage, [127]
- Catastrophes, geological, [145], [147]
- Caterpillars, food, [126], [127]
- Characters, acquired, inheritance of, [1], [57], [60], [225];
- Cirripedes, [201], [229]
- Classification, natural system of, [35], [199], [206], [208];
- Compensation, law of, [106]
- Conditions, direct, action of, [1], [57] n., [62], [65];
- Continent originating as archipelago, bearing of on distribution, [189]
- Cordillera, as channel of migration, [34] n., [191]
- Correlation, [76]
- Creation, centres of, [168], [192]
- Crocodile, [146]
- Cross-and Self-Fertilisation, early statement of principles of, [15], [69] n., [103] n.
- Crossing, swamping effect of, [2], [69], [96];
- Death, feigned by insects, [123]
- Difficulties, on theory of evolution, [15], [121], [128], [134]
- Disease, hereditary, [43] n., [58], [222]
- Distribution, geographical, [29], [31], [151], [174], [177];
- Disuse, inherited effects of, [46], [57]
- Divergence, principle of, [xxv], [37] n., [145] n., [208] n.
- Domestication, variation under, [57], [62];
- Ears, drooping, [236]
- Elevation, geological, favouring birth of new species, [32], [34] n., [35] n., [185]-[189];
- Embryo, branchial arches of, [42], [220];
- Embryology, [42], [218];
- Ephemera, selection falls on larva, [87] n.
- compared with that of 1842 and with the Origin, [xxii]
- Epizoa, [219]
- Essay of 1842, question as to date of, [xvi];
- Essay of 1844, writing of, [xvi];
- Evolution, theory of, why do we tend to reject it, [248]
- Expression, inheritance of, [114]
- Extinction, [23], [147], [192];
- Extinction and rarity, [198]
- Eye, [111] n., [128], [129], [130]
- Faculty, in relation to instinct, [123]
- Faunas, alpine, [30], [170], [188];
- Fauna and flora, of islands related to nearest land, [187]
- Fear of man, inherited, [17], [113]
- Fertility, interracial, [103], [104]
- Fish, colours of, [130], [131];
- Floras, alpine, [162];
- Flower, morphology of, [39], [216];
- Fly, causing extinction, [149]
- Flying, evolution of, [16], [131]
- Food, causing variations, [1], [58], [77], [78]
- Formation (geological) evidence from Tertiary system, [144];
- Formations, most ancient escape denudation in conditions unfavourable to life, [25], [139]
- Forms, transitional, [24], [35] n., [136], [142], [194];
- Fossils, Silurian, not those which first existed in the world, [26], [138];
- Fruit, attractive to animals, [130]
- Galapagos Islands and Darwin’s views, [xiv];
- Galapagos Islands, fauna, [31] n., [82]
- Gasteropods, embryology, [218]
- Genera, crosses between, [11], [97];
- Geography, in relation to geology, [31] n., [174], [177]
- Geographical distribution, see Distribution
- Geology, as producing changed conditions, [31];
- Glacial period, effect of on distribution of alpine and arctic plants, [165]
- Habit in relation to instinct, [17], [113], [115], [116]
- Habits in animals taught by parent, [18]
- Heredity, see Inheritance
- Homology of limbs, [38], [214]
- Homology, serial, [39], [215]
- Hybrid, fowls and grouse, [11];
- Hybrids, gradation in sterility of, [11], [72], [97];
- Individual, meaning of term, [58]
- and faculty, [18], [123];
- guided by reason, [18], [19], [118];
- migratory, [19];
- migratory, loss of by woodcocks, [120];
- migratory, origin of, [125];
- due to germinal variation rather than habit, [116];
- requiring education for perfection, [117];
- characterised by ignorance of end: e.g. butterflies laying eggs, [17], [118];
- butterflies laying eggs on proper plant, [118], [127];
- instinct, natural selection applicable to, [19], [120]
- Inheritance of acquired characters, see Character
- Inheritance, delayed or latent, [43], [44] n., [223];
- Insect, adapted to fertilise flowers, [87];
- Instinct, variation in, [17], [112];
- Instinct, for finding the way, [124];
- Intermediate forms, see Forms
- Island, see Elevation, Fauna, Flora
- Island, upheaved and gradually colonised, [184]
- Islands, nurseries of new species, [33], [35] n., [185], [189]
- Isolation, [32], [34] n., [64], [95], [183], [184]
- Antelope, [148];
- Armadillo, [174];
- Ass, [79], [107], [172];
- Bat, [38], [123], [128] n., [131], [132], [214];
- Bear, sterile in captivity, [100];
- —whale-like habit, [128] n.;
- Bizcacha, [168], [203], [212];
- Bull, mammæ of, [232];
- Carnivora, law of compensation in, [106];
- Cats, run wild at Ascension, [172];
- —tailless, [60];
- Cattle, horns of, [75], [207];
- Cheetah, sterility of, [100] and n.;
- Chironectes, [199];
- Cow, abortive mammæ, [232];
- Ctenomys, see Tuco-tuco;
- Dog, [106], [114];
- —in Cuba, [113] and n.;
- —mongrel breed in oceanic islands, [70];
- —difference in size a bar to crossing, [97];
- —domestic, parentage of, [71], [72], [73];
- —drooping ears, [236];
- —effects of selection, [66];
- —inter-fertile, [14];
- —long-legged breed produced to catch hares, [9], [10], [91], [92];
- —of savages, [67];
- —races of resembling genera, [106], [204];
- —Australian, change of colour in, [61];
- —bloodhound, Cuban, [204];
- —bull-dog, [113];
- —foxhound, [114], [116];
- —greyhound and bull-dog, young of resembling each other, [43], [44] n., [225];
- —pointer, [114], [115], [116], [117], [118];
- —retriever, [118] n.;
- —setter, [114];
- —shepherd-dog and harrier crossed, instinct of, [118], [119];
- —tailless, [60];
- —turnspit, [66];
- Echidna, [82] n.;
- Edentata, fossil and living in S. America, [174];
- Elephant, sterility of, [12], [100];
- Elk, [125];
- Ferret, fertility of, [12], [102];
- Fox, [82], [173], [181];
- Galeopithecus, [131] n.;
- Giraffe, fossil, [177];
- —tail, [128] n.;
- Goat, run wild at Tahiti, [172];
- Guanaco, [175];
- Guinea-pig, [69];
- Hare, S. American, [158] n.;
- Hedgehog, [82] n.;
- Horse, [67], [113], [115], [148], [149];
- Hyena, fossil, [177];
- Jaguar, catching fish, [132];
- Lemur, flying, [131] n.;
- Macrauchenia, [137];
- Marsupials, fossil in Europe, [175] n., [177];
- Mastodon, [177];
- Mouse, [153], [155];
- Mule, occasionally breeding, [97], [102];
- Musk-deer, fossil, [177];
- Mustela vison, [128] n., [132] n.;
- Mydas, [170];
- Mydaus, [170];
- Nutria, see Otter;
- Otter, [131], [132], [170];
- Pachydermata, [137];
- Phascolomys, [203], [212];
- Pig, [115], [217];
- Pole-cat, aquatic, [128] n., [132] _n._;
- Porpoise, paddle of, [38], [214];
- Rabbit, [74], [113], [236];
- Rat, Norway, [153];
- Reindeer, [125];
- Rhinoceros, [148];
- Ruminantia, [137] and n.;
- Seal, [93] n., [131];
- Sheep, [68], [78], [117], [205];
- Squirrel, flying, [131];
- Tapir, [135], [136];
- Tuco-tuco, blindness of, [46], [236];
- Whale, rudimentary teeth, [45], [229];
- Wolf, [71], [72], [82];
- Yak, [72]
- Lepidosiren, [140] n., [212]
- Limbs, vertebrate, of one type, [38], [216]
- Mammals, arctic, transported by icebergs, [170];
- Mammals, Names of:—
- Metamorphosis, literal not metaphorical, [41], [72]
- Metamorphosis, e.g. leaves into petals, [215]
- Migrants to new land, struggle among, [33], [185]
- Migration, taking the place of variation, [188]
- Monstrosities, as starting-points of breeds, [49], [59];
- Morphology, [38], [215];
- Mutation, see Sports
- extinct related to, existing in the same manner as representative existing ones to each other, [33], [192];
- introduced, beating indigenes, [153];
- dependent on other organisms rather than on physical surroundings, [185];
- graduated complexity in the great classes, [227];
- immature, how subject to natural selection, [42], [220], [228];
- all descended from a few parent-forms, [52], [252]
- Natural selection, see Selection
- Nest, bird’s, see Instinct
- Ocean, depth of, and fossils, [25], [195]
- Organisms, gradual introduction of new, [23], [144];
- Organs, perfect, objection to their evolution, [15], [128];
- Orthogenesis, [241] n.
- Oscillation of level in relation to continents, [33], [34] n., [241]
- Pallas, on parentage of domestic animals, [71]
- Ægilops, [58] n.;
- Artichoke (Jerusalem), [79];
- Ash, weeping, seeds of, [61];
- Asparagus, [79];
- Azalea, [13], [59], [97];
- Cabbage, [109], [135], [204];
- Calceolaria, [11], [99];
- Cardoon, [153];
- Carrot, variation of, [58] n.;
- Chrysanthemum, [59];
- Crinum, [11], [99];
- Crocus, [96], [99] n.;
- Cucubalus, crossing, [232];
- Dahlia, [21], [59], [63], [69], [74], [110];
- Foxglove, [82];
- Gentian, colour of flower, [107] n.;
- Geranium, [102];
- Gladiolus, crossed, ancestry of, [11];
- Grass, abortive flowers, [233];
- Heath, sterility, [96];
- Hyacinth, colours of, [106];
- —feather-hyacinth, [229];
- Juniperus, hybridised, [97];
- Laburnum, peculiar hybrid, [108];
- Lilac, sterility of, [13], [100];
- Marigold, style of, [47], [233], [237];
- Mistletoe, [6], [86], [87], [90] n.;
- Nectarines on peach trees, [59];
- Oxalis, colour of flowers of, [107] n.;
- Phaseolus, cultivated form suffers from frost, [107];
- Pine-apple, [207];
- Poppy, Mexican, [154];
- Potato, [69], [74], [110];
- Rhododendron, [97], [99];
- Rose, moss, [59];
- —Scotch, [69];
- Seakale, [79];
- Sweet-william, [59];
- Syringa, persica and chinensis, see Lilac;
- Teazle, [129];
- Thuja, hybridised, [97];
- Tulips, "breaking" of, [58];
- Turnip, Swedish and common, [205];
- Vine, peculiar hybrid, [108];
- Yew, weeping, seeds of, [61]
- Pampas, imaginary case of farmer on, [32], [184]
- Perfection, no inherent tendency towards, [227]
- Plants, see also Flora;
- Plants, Names of:—
- Plasticity, produced by domestication, [1], [63]
- Plesiosaurus, loss of unity of type in, [41], [217]
- Pteropods, embryology, [218]
- Quadrupeds, extinction of large, [147]
- Quinary System, [202]
- Race, the word used as equivalent to variety, [94]
- Races, domestic, classification of, [204]
- formerly less distinct as judged by fossils, [177]
- Rarity, [28], [148];
- Recapitulation theory, [42], [219], [230], [239]
- Record, geological, imperfection of, [26], [140]
- Regions, geographical, of the world, [29], [152], [174];
- Resemblance, analogical, [36], [199]
- Reversion, [3], [64], [69], [74]
- “Roguing,” [3]
- Rudimentary organs, see Organs
- Savages, domestic animals of, [67], [68], [96]
- Selection, human, [3], [63];
- Selection, natural, [xvi], [7], [87];
- Selection, sexual, two types of, [10], [92]
- Silk-worms, variation in larval state, [44] n., [223]
- Skull, morphology of, [39], [215]
- Species, representative, seen in going from N. to S. in a continent, [31] n., [156];
- Species, not created more than once, [168], [171], [191];
- Sports, [1], [58], [59], [64], [74], [95], [129], [186], [206], [224]
- Sterility, due to captivity, [12], [77] n., [100];
- Struggle for life, [7], [91], [92], [148], [241]
- Subsidence, importance of, in relation to fossils, [25], [35] n., [7];
- Swimming bladder, [16], [129]
- System, natural, is genealogical, [36], [208]
- Telegony, [108]
- Tibia and fibula, [48], [137]
- Time, enormous lapse of, in geological epochs, [25], [140]
- Tortoise, [146]
- Transitional forms, see Forms
- Trigonia, [147] n., [199]
- Tree-frogs in treeless regions, [131]
- produced by change and also by crossing, [105]
- Type, unity of, [38], [214];
- under domestication, [1], [57], [63], [78];
- due to causes acting on reproductive system, see Variation, germinal;
- individual, [57] n.;
- causes of, [1], [4], [57], [61];
- due to crossing, [68], [69];
- limits of, [74], [75], [82], [109];
- small in state of nature, [4], [59] n., [81], [83];
- results of without selection, [84];
- —minute, value of, [91];
- analogous in species of same genus, [107];
- of mental attributes, [17], [112];
- in mature life, [59], [224], [225]
- Uniformitarian views of Lyell, bearing on evolution, [249]
- Use, inherited effects of, see Characters, acquired
- Variability, as specific character, [83];
- Variation, by Sports, see Sports;
- Varieties, minute, in birds, [82];
- Vertebrate skull, morphology of, [215]
- Wildness, hereditary, [113], [119]
Footnotes
[{1}] See the extracts in Life and Letters of Charles Darwin, ii. p. 5.
[{2}] The second volume,—especially important in regard to Evolution,—reached him in the autumn of 1832, as Prof. Judd has pointed out in his most interesting paper in Darwin and Modern Science. Cambridge, 1909.
[{3}] Obituary Notice of C. Darwin, Proc. R. Soc. vol. 44. Reprinted in Huxley's Collected Essays. See also Life and Letters of C. Darwin, ii. p. 179.