In a highly organised and complex animal, the gemmules thrown off from each different cell or unit throughout the body must be inconceivably numerous and minute. Each unit of each part, as it changes during development, and we know that some insects undergo at least twenty metamorphoses, must throw off its gemmules. All organic beings, moreover, include many dormant gemmules derived from their grandparents and more remote progenitors, but not from all their progenitors. These almost infinitely numerous and minute gemmules must be included in each bud, ovule, spermatozoon, and pollen-grain. Such an admission will be declared impossible; but, as previously
remarked, number and size are only relative difficulties, and the eggs or seeds produced by certain animals or plants are so numerous that they cannot be grasped by the intellect.
The organic particles with which the wind is tainted over miles of space by certain offensive animals must be infinitely minute and numerous; yet they strongly affect the olfactory nerves. An analogy more appropriate is afforded by the contagious particles of certain diseases, which are so minute that they float in the atmosphere and adhere to smooth paper; yet we know how largely they increase within the human body, and how powerfully they act. Independent organisms exist which are barely visible under the highest powers of our recently-improved microscopes, and which probably are fully as large as the cells or units in one of the higher animals; yet these organisms no doubt reproduce themselves by germs of extreme minuteness, relatively to their own minute size. Hence the difficulty, which at first appears insurmountable, of believing in the existence of gemmules so numerous and so small as they must be according to our hypothesis, has really little weight.
The cells or units of the body are generally admitted by physiologists to be autonomous, like the buds on a tree, but in a less degree. I go one step further and assume that they throw off reproductive gemmules. Thus an animal does not, as a whole, generate its kind through the sole agency of the reproductive system, but each separate cell generates its kind. It has often been said by naturalists that each cell of a plant has the actual or potential capacity of reproducing the whole plant; but it has this power only in virtue of containing gemmules derived from every part. If our hypothesis be provisionally accepted, we must look at all the forms of asexual reproduction, whether occurring at maturity or as in the case of alternate generation during youth, as fundamentally the same, and dependent on the mutual aggregation and multiplication of the gemmules. The regrowth of an amputated limb or the healing of a wound is the same process partially carried out. Sexual generation differs in some important respects, chiefly, as it would appear, in an insufficient number of gemmules being aggregated within the separate sexual elements, and probably in the presence of certain primordial cells. The development of each being, including all the
forms of metamorphosis and metagenesis, as well as the so-called growth of the higher animals, in which structure changes though not in a striking manner, depends on the presence of gemmules thrown off at each period of life, and on their development, at a corresponding period, in union with preceding cells. Such cells may be said to be fertilised by the gemmules which come next in the order of development. Thus the ordinary act of impregnation and the development of each being are closely analogous processes. The child, strictly speaking, does not grow into the man, but includes germs which slowly and successively become developed and form the man. In the child, as well as in the adult, each part generates the same part for the next generation. Inheritance must be looked at as merely a form of growth, like the self-division of a lowly-organised unicellular plant. Reversion depends on the transmission from the forefather to his descendants of dormant gemmules, which occasionally become developed under certain known or unknown conditions. Each animal and plant may be compared to a bed of mould full of seeds, most of which soon germinate, some lie for a period dormant, whilst others perish. When we hear it said that a man carries in his constitution the seeds of an inherited disease, there is much literal truth in the expression. Finally, the power of propagation possessed by each separate cell, using the term in its largest sense, determines the reproduction, the variability, the development and renovation of each living organism. No other attempt, as far as I am aware, has been made, imperfect as this confessedly is, to connect under one point of view these several grand classes of facts. We cannot fathom the marvellous complexity of an organic being; but on the hypothesis here advanced this complexity is much increased. Each living creature must be looked at as a microcosm—a little universe, formed of a host of self-propagating organisms, inconceivably minute and as numerous as the stars in heaven.
CHAPTER XXVIII.
CONCLUDING REMARKS.
DOMESTICATION—NATURE AND CAUSES OF VARIABILITY—SELECTION—DIVERGENCE AND DISTINCTNESS OF CHARACTER—EXTINCTION OF RACES—CIRCUMSTANCES FAVOURABLE TO SELECTION BY MAN—ANTIQUITY OF CERTAIN RACES—THE QUESTION WHETHER EACH PARTICULAR VARIATION HAS BEEN SPECIALLY PREORDAINED.