perfection had been attained with our flowers, but a higher standard has soon been reached. Hardly any fruit has been more improved than the strawberry, yet a great authority remarks,[[597]] "it must not be concealed that we are far from the extreme limits at which we may arrive."
Time is an important element in the formation of our domestic races, as it permits innumerable individuals to be born, and these when exposed to diversified conditions are rendered variable. Methodical selection has been occasionally practised from an ancient period to the present day, even by semi-civilised people, and during former times will have produced some effect. Unconscious selection will have been still more effective; for during a lengthened period the more valuable individual animals will occasionally have been saved, and the less valuable neglected. In the course, also, of time, different varieties, especially in the less civilised countries, will have been more or less modified through natural selection. It is generally believed, though on this head we have little or no evidence, that new characters in time become fixed; and after having long remained fixed it seems possible that under new conditions they might again be rendered variable.
How great the lapse of time has been since man first domesticated animals and cultivated plants, we begin dimly to see. When the lake-buildings of Switzerland were inhabited during the Neolithic period, several animals were already domesticated and various plants cultivated. If we may judge from what we now see of the habits of savages, it is probable that the men of the earlier Stone period—when many great quadrupeds were living which are now extinct, and when the face of the country was widely different from what it now is—possessed at least some few domesticated animals, although their remains have not as yet been discovered. If the science of language can be trusted, the art of ploughing and sowing the land was followed, and the chief animals had been already domesticated, at an epoch so immensely remote, that the Sanskrit, Greek, Latin, Gothic, Celtic, and Sclavonic languages had not as yet diverged from their common parent-tongue.[[598]]
It is scarcely possible to overrate the effects of selection occasionally carried on in various ways and places during thousands of generations. All that we know, and, in a still stronger degree, all that we do not know,[[599]] of the history of the great majority of our breeds, even of our more modern breeds, agrees with the view that their production, through the action of unconscious and methodical selection, has been almost insensibly slow. When a man attends rather more closely than is usual to the breeding of his animals, he is almost sure to improve them to a slight extent. They are in consequence valued in his immediate neighbourhood, and are bred by others; and their characteristic features, whatever these may be, will then slowly but steadily be increased, sometimes by methodical and almost always by unconscious selection. At last a strain, deserving to be called a sub-variety, becomes a little more widely known, receives a local name, and spreads. The spreading will have been extremely slow during ancient and less civilised times, but now is rapid. By the time that the new breed had assumed a somewhat distinct character, its history, hardly noticed at the time, will have been completely forgotten; for, as Low remarks,[[600]] "we know how quickly the memory of such events is effaced."
As soon as a new breed is thus formed, it is liable through the same process to break up into new strains and sub-varieties. For different varieties are suited for, and are valued under, different circumstances. Fashion changes, but, should a fashion last for even a moderate length of time, so strong is the principle of inheritance, that some effect will probably be impressed on the breed. Thus varieties go on increasing in number, and history shows us how wonderfully they have increased since the earliest records.[[601]] As each new variety is produced, the earlier, intermediate, and less valuable forms will be neglected, and perish. When a breed, from not being valued, is kept in small numbers, its extinction almost inevitably follows sooner or later, either from accidental causes of destruction or from close interbreeding; and this is an event which, in the case of well-marked breeds, excites attention. The birth or production of a new domestic race is so slow a process that it
escapes notice; its death or destruction is comparatively sudden, is often recorded, and when too late sometimes regretted.
Several authors have drawn a wide distinction between artificial and natural races. The latter are more uniform in character, possessing in a high degree the character of natural species, and are of ancient origin. They are generally found in less civilised countries, and have probably been largely modified by natural selection, and only to a small extent by man's unconscious and methodical selection. They have, also, during a long period, been directly acted on by the physical conditions of the countries which they inhabit. The so-called artificial races, on the other hand, are not so uniform in character; some have a semi-monstrous character, such as "the wry-legged terriers so useful in rabbit-shooting,"[[602]] turnspit dogs, ancon sheep, niata oxen, Polish fowls, fantail-pigeons, &c.; their characteristic features have generally been acquired suddenly, though subsequently increased in many cases by careful selection. Other races, which certainly must be called artificial, for they have been largely modified by methodical selection and by crossing, as the English race-horse, terrier-dogs, the English game-cock, Antwerp carrier-pigeons, &c., nevertheless cannot be said to have an unnatural appearance; and no distinct line, as it seems to me, can be drawn between natural and artificial races.
It is not surprising that domestic races should generally present a different aspect from natural species. Man selects and propagates modifications solely for his own use or fancy, and not for the creature's own good. His attention is struck by strongly marked modifications, which have appeared suddenly, due to some great disturbing cause in the organisation. He attends almost exclusively to external characters; and when he succeeds in modifying internal organs,—when for instance he reduces the bones and offal, or loads the viscera with fat, or gives early maturity, &c.,—the chances are strong that he will at the same time weaken the constitution. On the other hand, when an animal has to struggle throughout its life with many competitors and enemies, under circumstances inconceivably complex and liable to change, modifications of the most varied nature—in the internal organs as well as in external characters, in the
functions and mutual relations of parts—will be rigorously tested, preserved, or rejected. Natural selection often checks man's comparatively feeble and capricious attempts at improvement; and if this were not so, the result of his work, and of nature's work, would be even still more different. Nevertheless, we must not overrate the amount of difference between natural species and domestic races; the most experienced naturalists have often disputed whether the latter are descended from one or from several aboriginal stocks, and this clearly shows that there is no palpable difference between species and races.